Book IV

© Brian W. Coad (www.briancoad.com)

Book I = Introduction to Chanidae

Book II = Cyprinidae

Book III = Cobitidae to Marine Species

Book IV = Checklists to Bibliography

E. Checklists

Farsi Names

?add names from Species Accounts

The purpose of this list is to bring together the Farsi, English and scientific names of the freshwater fishes of Iran. Confusion over the names of fishes can render any study or article confusing or even useless. Generally the best course to follow is to use the scientific name, at least initially, and then use the common name. However, common names are often more familiar or evocative to readers, especially in popular and newspaper articles.

Effective management of resources as well as scientific and popular studies and publications need a clear link between common and scientific names. Such linkages also facilitate access to the extensive world literature on fishes and their biology.

This is by no means a final listing; I hope to elicit a response from readers of this section in order to add common names in Farsi to the list.

Farsi Names

The Farsi name(s) are given below the scientific name. They have been obtained from literature sources (listed in Coad (1995; 1999)) but primarily in discussions with Iranian ichthyologists. These names may have a meaning if translated into English but since these are never or very rarely used they are not translated here (translations can be found in the Species Accounts). A prime example would be safid mahi (= white fish) which occasionally appears in travel and scientific literature under this English name. Confusion has arisen over the use of this English translation since whitefish in English scientific literature refers to members of the salmon family (Salmonidae, usually the genus Coregonus) and not this Iranian member of the carp family (Cyprinidae, Rutilus frisii).

Some of the names are highly original and evocative in English translation while others are direct translations of the English names. Interesting names include ordak mahi (= duck fish in reference to the characteristic snout of the pike), gel-cheragh (= mud-eater or mud-grazer in relation to feeding habits for several unrelated fishes), gour-e khar (= striped donkey, i.e. zebra, for Aphanius spp.) and heif-e noon (= waste of bread, i.e. valueless, used in Khuzestan for Chondrostoma regium).

A number of Iranian species have no common Farsi name that I have been able to discover and are indicated by a question mark (?). Doubtful names are also indicated by a question mark. Any reader who can supply a common name and is sure of the species identification is asked to email me (see www.briancoad.com for contact methods). Ideally a preserved specimen of the fish should be sent to me for verification to avoid confusion and misunderstandings.

Some genera and families have a common name without a unique modifier for each species. These names are included here. I have not added modifiers in case a reader informs me of a common name. In future, if no common name seems to exist in Farsi, then it may be advisable to invent one to separate the species.

Several names exist for certain species, used locally. Some common name lists standardise a common name for each species but I have not attempted this here. More research and feedback from readers is needed to determine which names are the most commonly used, widespread and acceptable before one is chosen in preference to another. The variety of names is a regional heritage and should not be casually ignored. Robins et al. (1991a) give reasons and criteria for establishing a single common name for each species.

English Names

The English name(s) in commonest or official use are given here in parentheses below the Farsi name. Some Iranian species lack English names for the simple reason that these fishes do not occur in English-speaking countries nor are they widely known as aquarium, food or sport fishes. A single English name has not been selected as the primary aim of this work is to link scientific and Farsi names and secondarily to indicate the English names of the species as an aid in data retrieval and comprehension. Note that many English names are composed of a noun and a modifying adjective. For simplicity the noun is not repeated each time with its adjective so there is "common, Oriental, eastern, bronze or carp bream" rather than "common bream, Oriental bream, eastern bream, bronze bream and carp bream". A semicolon (;) may separate words which are not part of a series of adjectives. Thus "beluga; great, giant or European sturgeon" indicates that "beluga" is a fish name by itself while sturgeon may have the adjectives great, giant and European.

Reference sources include Robins et al. (1991a; 1991b) for North American native species and species important to North Americans, Berg (1948-1949), Lindberg and Heard (1972) and Ricker (1973) for the former U.S.S.R. and adjacent countries, and OECD (1995) for official names used by the Food and Agriculture Organization of the United Nations. For Caspian Sea basin species, the name is often taken from the common usage in Berg (1948-1949) where the Russian name has been given its translation. Berg's work has been widely used as a reference source on fishes in the Caspian Sea and adjacent basins since the original Russian work is also available in English translation. Note that some Russian names have no direct equivalent in English and the Cyrillic orthography is converted to English. Purely Russian names are not included when well-known English names are available, e.g. the Russian "nalim" for burbot or eelpout (Lota lota). Some Russian names are in common use in the Caspian basin of Iran even though there are Farsi names for the fishes. The northern fisheries were once controlled by Russian experts. An example of this is Huso huso, the beluga (a Russian word but also used in Farsi and English) or fil mahi (purely Farsi). Alternative names, such as great or giant sturgeon, do not appear commonly in Iranian scientific and popular works despite being more descriptive and evocative in English and having the advantage of not being confused with the whale of the same name. Similarly, kutum (safid mahi, Rutilus frisii kutum) is often used in scientific literature as a common name and is included here although there is a less well-known English version (Southern Caspian roach).

An exhaustive listing of Russian, Turkmen and Azerbaijani names is not given here and the reader is referred to the Russian edition of Berg (1948-1949) and Lindberg and Heard (1972) for further details. Some names in Gilaki and Mazanderani from the Caspian coast of Iran have been given from sources in the Encyclopaedia Iranica, Columbia University Press, New York (?website). Some names from Turkmenistan and Azerbaijan may be used in Iran, especially near the borders, but I have no information on this usage. Similarly Heckel (1843, 1846-1849) lists a number of Arabic names of fishes which also occur in Iran, although these works are now probably out of date. More recent common names can be found in Khalaf (1961), Mahdi (1962), Al-Nasiri and Hoda (1976) and other works on Iraqi fishes (see Coad and Al-Hassan (1988) for a bibliography on these fishes). Arabic common names are used in Khuzestan and are included here, modified in Farsi, where this usage is known.

A summary of works on Turkish fishes is found in Coad and Kuru (1986). Fishes of Pakistan are summarised in Mirza (1972, 1990) with a bibliography on this fauna in Mirza (1978). I am not aware if common names from Pakistan are used widely in Iranian Baluchestan and so these are not included here. The English common name presents problems where several subspecies, "morpha" or "forms" have been described within a species. This is almost entirely a problem of the Caspian Sea basin where Russian workers have distinguished many subspecies of the families Clupeidae and Cyprinidae in particular. Each of these has a common name, at least in Russian, usually relating to the locality of the subspecies. Some recent studies have demonstrated that these subspecies are often ill-founded and even distinct species have been shown to be the same as more widespread ones outside the Caspian Sea basin (Coad, 1996b). I have listed all the available names in English including those with adjectives which seem to be geographically inappropriate. For example, Barbus brachycephalus is called the Aral barbel in Robins et al. (1991b), presumably based on the type subspecies found in the Aral Sea basin. A distinct subspecies in the Caspian Sea basin, Barbus brachycephalus caspius, is more appropriately called the Caspian barbel here. Furthermore, many of these Russian common names for subspecies and their English translations are "book names" - artificial constructs not used by local people. An additional problem lies with nomenclatural changes in the scientific name. Alosa pontica, a member of the family Clupeidae, was previously thought to inhabit only the waters of the Black Sea and was appropriately named in English as the Pontic shad or Black Sea herring. As now understood this species is also found in the Caspian Sea (albeit as a distinct subspecies) and the English name is inappropriate. In such cases I have also retained the older name for the Caspian Sea species or subspecies. Conversely when a species or subspecies previously known only from, for example, the Caspian Sea has been shown to be synonymous with a more widespread species then the name of the widespread species is added to the list. An example is the short-snouted pipefish, Syngnathus abaster, once regarded as a distinct subspecies in another species (Syngnathus nigrolineatus caspius or Caspian pipefish).

Scientific Names

The most recent scientific name is given for each species. It should be understood that scientific names are dynamic, changing as our understanding of the relationships of fishes changes. A recent reference on the scientific names and nomenclatorial problems of Iranian freshwater fishes is Coad (1995). The current list also serves as an updated version of that work. Changes include the placement of Pseudophoxinus persidis in the genus Leuciscus (after N. Bogutskaya, pers. comm., 1995) and the distinction of Aphanius persicus (Coad, 1996a). Lazara (1995) proposes the use of the genus Lebias over Aphanius in Cyprinodontidae but Kottelat (1997) disagrees. Kottelat (1997) also demonstrates that Sander has priority over Stizostedion and along with Eschmeyer (1998) lists other minor changes. Elvira (1997) resurrects Chondrostoma cyri. Nalbant and Bianco (1998) describe 5 new species of Balitoridae from Iran. The arrangement of families and orders also changes with time and this too is reflected here. The arrangement has been changed from that used in Coad (1995) to accord with Nelson (1994), which was not then available.

Coverage

A brief listing of distribution is given for each species after the scientific name based on drainage basins (see Coad (1995) for map). The limits of this article are the political boundaries of Iran including the Caspian Sea south of a line drawn between the borders of Iran and Azerbaijan in the west and Iran and Turkmenistan in the east. The Caspian Sea has a salt concentration about one-third seawater and the fauna there is a mixture of freshwater and brackish or marine species. A few species in the south are included although they are thought of as primarily marine. An example is a single shark species which can reside in fresh waters and has considerable importance since it preys on humans (Coad and Papahn, 1988). The scientific name list contains more marine species that may enter fresh waters.

Format

Classes, orders and families are listed in phylogenetic sequence following Nelson (1994). Within families species are listed alphabetically by scientific name. A separate alphabetical list of Farsi names is appended for convenience.

Class 1. Cephalaspidomorphi

Order 1. Petromyzontiformes

Family 1. Petromyzontidae (lampreys)

1. Caspiomyzon wagneri (Kessler, 1870) - Caspian Sea basin.

mar mahi, mar mahi dahangerd, mahi dahangerd daryacheh-ye khazar.

(Caspian or Volga lamprey).

Class 2. Chondrichthyes

Order 2. Carcharhiniformes

Family 2. Carcharhinidae (requiem sharks)

1. Carcharhinus leucas (Valenciennes in Müller and Henle, 1839) - Tigris River basin.

kooseh, sag mahi.

(bull shark).

Class 3. Actinopterygii

Order 3. Acipenseriformes

Family 3. Acipenseridae (sturgeons)

1. Acipenser gueldenstaedtii Brandt in Brandt and Ratzeburg, 1833 - Caspian Sea basin.

chalbash, tass mahi (this term includes A. gueldenstaedtii, and A. nudiventris for large eggs, in fisheries statistics), تاس ماهي روس (= tass mahi Rus), osetra, sag mahi, mahi-ye kaviar, kaviari rusi.

(osetr; Russian, Kura, Persian or Azov-Black Sea sturgeon).

2. Acipenser nudiventris Lovetzky, 1828 - Caspian Sea basin.

ship, šep, tass mahi (included under this name with A. gueldenstaedtii and A. persicus when eggs are large for fisheries statistics), tass mahi shekam brahne, sag mahi, mahi-ye kaviar, keshdi, shenavar.

(spiny, thorn or fringebarbel sturgeon; ship or sheap).

3. Acipenser persicus Borodin, 1897 - Caspian Sea basin.

kara burun, darakul, tirij, tass mahi (this term includes A. gueldenstaedtii), tass mahi Iran, tas mahi Irani, sag mahi, mahi-ye kaviar.

(Persian or Kura sturgeon).

4. Acipenser stellatus Pallas, 1771 - Caspian Sea basin.

uzun burun, darakul, sevruga, sevroga, sag mahi, mahi-ye kaviar, puze draz.

(sevruga or stellate sturgeon (this term also includes A. nudiventris with small eggs for fisheries statistics), South Caspian stellate sturgeon, star sturgeon).

5. Huso huso (Linnaeus, 1758) - Caspian Sea basin.

fil mahi, beluga, beloga, mahi kaviar-e bozorg, sag mahi, mahi-ye kaviar.

(beluga; great, giant or European sturgeon).

Order 4. Anguilliformes

Family 4. Anguillidae (freshwater eels)

1. Anguilla anguilla (Linnaeus, 1758) - Caspian Sea basin.

mar mahi ma'muli.

(European, common, silver or yellow eel).

Order 5. Clupeiformes

Family 5. Clupeidae (herrings)

1. Alosa braschnikowii (Borodin, 1904) - Caspian Sea basin.

shag mahi, shag mahi-ye Khazari.

(Caspian marine shad; Kura, striped, Agrakhan, bigeye, dolginka, whitehead, Astrabad, yellow-back, Gasan-Kuli, Kiselevitch, Krasnovodsk, eastern, Sara, or May herring; driver, glutton).

2. Alosa caspia (Eichwald, 1838) - Caspian Sea basin.

shag mahi shekam bozorg, shag mahi chekameh dar, shag mahi darya-ye khazar, zalun (in Gilaki), puzanok.

(Caspian, North Caspian, Central Caspian, il'men, Enzeli, Sara, Baku or Astrabad shad).

3. Alosa pontica (Eichwald, 1838) - Caspian Sea basin.

shag mahi-ye posht siah, shag mahi darya-ye siah, zalun (in Gilaki), puzanok.

(blackback, blacknose, Caspian anadromous shad; black-spined, Volga, many-rakered or Astrakhan herring; Pontic shad, Black Sea herring).

4. Alosa saposchnikowii (Grimm, 1887) - Caspian Sea basin.

shag mahi-e chesm dorosht, shag mahi.

(bigeye or Saposhnikovi shad).

5. Alosa sphaerocephala (Berg, 1913) - Caspian Sea basin.

shag mahi Agrakhan.

(roundheaded or Agrakhan shad).

6. Clupeonella cultriventris (Nordmann, 1840) - Caspian Sea basin.

kilka or kelka, rizeh keraye, rizeh kuli, kilka-ye ma'muli.

(common kilka or tyulka; sardelle; Caspian sprat, kilka or tyulka; common Caspian kilka, Black Sea sprat).

7. Clupeonella engrauliformis (Borodin, 1904) - Caspian Sea basin.

kilka or kelka, rizeh keraye, kilka-ye anchovy.

(anchovy sprat or kilka, anchovy-like tyulka, sardelle).

8. Clupeonella grimmi Kessler, 1877 - Caspian Sea basin.

kilka or kelka, kilka-ye cheshm dorosht.

(South Caspian sprat, bigeye kilka or tyulka, sardelle).

9. Tenualosa ilisha (Hamilton-Buchanan, 1822) - Tigris River basin; probably other rivers of southern Iran.

sobur, zobur, zabur, zomur, brk, mahi khor kuchiku.

(hilsa; Indian, river or hilsa shad; ilisha).

Order 6. Gonorhynchiformes

Family 6. Chanidae (milkfishes)

1. Chanos chanos (Forsskål, 1775) - Hormuz basin.

khameh mahi.

(milkfish, salmon herring)

Order 7. Cypriniformes

Family 7. Cyprinidae (carps and minnows)

1. Abramis brama (Linnaeus, 1758) - Caspian Sea basin.

sim, mahi-ye sim.

(common, Oriental, eastern, bronze or carp bream).

2. Abramis sapa (Pallas, 1814) - Caspian Sea basin.

sim, sapa, mahi sim kondpuzh.

(white-eye, South Caspian or southern white-eye bream).

3. Acanthalburnus microlepis (De Filippi, 1863) - Caspian Sea basin.

?

(blackbrow bleak).

4. Acanthalburnus urmianus (Günther, 1899) - Lake Orumiyeh basin.

?

(None).

5. Acanthobrama marmid Heckel, 1843 - Tigris River basin.

kalashpa, shebhe-sardin, shebhe-nazy.

(None).

6. Alburnoides bipunctatus (Bloch, 1782) - Caspian Sea, Lake Orumiyeh, Namak Lake, Esfahan, Kor, Tigris and Tedzhen (= Hari) River basins.

mahi khiateh or khayateh, parak, lapak (in Mazandaran), shebhe zury (in Khuzestan).

(riffle minnow or bleak; spirlin).

7. Alburnus alburnus (Linnaeus, 1758) - Caspian Sea basin; widely introduced.

mahi morvarid, kuli Irani, kuli-ye Irani.

(bleak, Transcaucasian, Persian or Safid River bleak).

8. Alburnus atropatenae Berg, 1925 - Lake Orumiyeh basin.

?

(None).

9. Alburnus chalcoides (Gueldenstaedt, 1772) - Caspian Sea basin.

shah kuli (in Gilaki), shah mahi; aslak (in Mazandaran); siah kuli, safid kuli.

(bleak, Caspian shemaya, Danube bleak).

10. Alburnus filippii Kessler, 1877 - Caspian Sea basin.

kuli Kura, kuli-ye Kura.

(Kura or Filippi's bleak).

11. Alburnus mossulensis Heckel, 1843 - Esfahan, Tigris and Kor Rivers, Gulf, Hormuz and Lake Maharlu basins.

shah kuli-ye jonubi.

(None).

12 Aspidoparia morar (Hamilton-Buchanan, 1822) - Makran and Mashkid River basins.

?

(None).

13. Aspius aspius (Linnaeus, 1758) - Caspian Sea basin.

mash mahi, khasham.

(asp; redlip, Caspian or South Caspian asp).

14. Aspius vorax Heckel, 1843 - Tigris River basin.

shelej, sholge.

(None).

15. Barbus barbulus Heckel, 1849 - Gulf, Tigris and Kor River basins.

lab-pahn; berzem or berzem lab-pahn (in Khuzestan); boz mahi or sos mahi (in the Dalaki and Shapur river basins).

(None).

16. Barbus brachycephalus Kessler, 1872 - Caspian Sea basin.

zardek, zardehpar, sos mahi, sas mahi khazari.

(Caspian barbel, Aral barbel)

17. Barbus capito (Gueldenstaedt, 1773) - Caspian Sea basin.

usach bulatmai, usach chanari, zardek, zard pareh, zardehpar, rnj, sos mahi, pulad mahi, ses mahi bozorg.

(None).

18. Barbus esocinus (Heckel, 1843) - Tigris River basin.

soong (in northern Khuzestan and Lorestan); anzeh and bach (at Ahvaz and in southern Khuzestan); bizz, narbach; balzard.

("Tigris salmon", "Euphrates salmon", pike barb).

19. Barbus grypus Heckel, 1843 - Tigris River, Gulf and Hormuz basins.

shirbot or shilbot, shaboot, rumi, sos mahi, shebhe shirbot; sorkheh (in the Zohreh River).

(large scaled barb).

20. Barbus kersin Heckel, 1843 - Gulf, Tigris and Kor River basins.

berzem.

(None).

21. Barbus kosswigi (Ladiges, 1960) - Tigris River basin.

abu henesh, shebhe shirbot.

(None).

22. Barbus lacerta Heckel, 1843 - Caspian Sea, Lake Orumiyeh and Tigris River basins.

blizem

(Kura barbel).

23. Barbus luteus (Heckel, 1843) - Tigris River, Kor River, Lake Maharlu, Gulf and Hormuz basins.

himri, sangal.

(golden barb).

24. Barbus mursa (Gueldenstaedt, 1773) - Caspian Sea and Namak Lake basins.

sos mahi, mahi siah, zardek-e qalami, ses mahi koloft safid rud.

(mursa).

25. Barbus pectoralis Heckel, 1843 - Gulf, Tigris and Kor River basins.

bassan, berzem, ? touyeni.

(None).

26. Barbus sharpeyi Günther, 1874 - Tigris River basin.

binni, soleimani.

(None).

27. Barbus sublimus Coad and Najafpour, 1997

?

(None).

28. Barbus subquincunciatus Günther, 1868 - Tigris River basin.

soleimani.

(black spot barb).

29. Barbus xanthopterus (Heckel, 1843) - Tigris River basin.

gatan.

(None).

30. Barilius mesopotamicus Berg, 1932 - Tigris River and Gulf basins.

?

(None).

31. Blicca bjoerkna (Linnaeus, 1758) - Caspian Sea basin.

simparek, mahi sim nama.

(Transcaucasian white bream; silver, white or flat bream).

32. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844) - Namak Lake, Kavir, Kerman-Na'in, Esfahan, Kor and Tigris River basins.

shum, siah mahi.

(None).

33. Capoeta barroisi (Lortet in Barrois, 1894) - Tigris River and Gulf basins.

siah mahi Dasht-e Arzhani.

(None).

34. Capoeta buhsei Kessler, 1877 - Namak Lake basin.

shamshiri; mahi sibeli (at Karaj).

(None).

35. Capoeta capoeta (Gueldenstaedt, 1773) - Caspian Sea, Tedzhen (= Hari) River, Lake Orumiyeh, Kavir, Bejestan, Sirjan, Namak Lake, Esfahan, Tigris River and Gulf basins.

tilkhos, siah mahi; soru (in the Dalaki and Shapur river basins)

(Lenkoran, Kura, or Transcaspian khramulya or barb).

36. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842) - Namak Lake, Esfahan, Tigris River, Gulf, Kor River, Lake Maharlu, Sirjan, Kerman-Na'in, Hormuz, Lut and Hamun-e Jaz Murian basins.

zardeh or siah mahi (in the Caspian basin); touyeni, gel-cheragh or gel khorok (in Khuzestan); qezel ala (in Chahar Mahall).

(None).

37. Capoeta fusca Nikolskii, 1897 - Tedzhen (= Hari) River, Bejestan, Sistan and Lut basins.

siah mahi.

(None).

38. Capoeta trutta (Heckel, 1843) - Tigris River basin.

touyeni (in Khuzestan); shir mahi, barg bidy, ? berzem.

(None).

39. Carassius auratus (Linnaeus, 1758) - Widely introduced.

mahi-ye talaee (in Khuzestan); ooshin; kopur-cheh or karaz (in Mazandaran); mahi-ye howz, mahi howz noqreai.

(silver crucian carp, goldfish, Prussian carp).

40. Chondrostoma cyri Kessler, 1877 - Caspian Sea basin.

?

(Kura nase; blackbelly; Kura undermouth).

41. Chondrostoma regium (Heckel, 1843) - Tigris and Kor Rivers and northern Gulf basins.

jokhorak, nazok; heif-e noon (in Khuzestan).

(None).

42. Crossocheilus latius (Hamilton-Buchanan, 1822) - Makran, Mashkid River and Sistan basins.

?

(None).

43. Ctenopharyngodon idella (Valenciennes in Cuvier and Valenciennes, 1844) - Widely introduced.

alaf khoar or kopur alaf khoar; amur (in Khuzestan).

(grass carp, white amur).

44. Cyprinion kais Heckel, 1843 - Tigris River and Gulf basins.

butak-e dehan kuchek, butok, lotak.

(None).

45. Cyprinion macrostomum Heckel, 1843 - Tigris River and Gulf basins.

botak, lotak or butak-e dehan buzorg (in Khuzestan); kopur, ? galuk.

(large-mouthed barb).

46. Cyprinion milesi (Day, 1880) - Hormuz, Hamun-e Jaz Murian and Makran basins.

?

(None).

47. Cyprinion tenuiradius Heckel, 1849 - Kor River, Lake Maharlu and Gulf basins.

?

(None).

48. Cyprinion watsoni (Day, 1872) - Hormuz, Hamun-e Jaz Murian, Mashkid River, Makran, Sistan and Lut basins.

?

(None).

49. Cyprinus carpio Linnaeus, 1758 - Tedzhen (= Hari) River and Caspian Sea basins. Widely introduced.

kopur, kapur (in Gilaki), kopur-e aineheh, kopur-e ma'muli; rashti or kopur-e rashti (in Khuzestan); mahi-ye gul.

(common, mirror, leather, European or German carp; koi).

50. Garra persica Berg, 1913 - Hamun-e Jaz Murian, Hormuz basins and Sistan basins.

gel-cheragh.

(None).

51. Garra rossica (Nikolsky, 1900) - Tedzhen (= Hari) River, Bejestan, Sistan, Lut, Hamun-e Jaz Murian, Mashkid River and Makran basins.

gel-cheragh, ? anjarak.

(None).

52. Garra rufa (Heckel, 1843) - Tigris River, Kor River, Lake Maharlu and Gulf basins, Hormuz basin.

shirbot, gel khorok, gel-cheragh, gararufa.

(None).

53. Garra variabilis (Heckel, 1843) - Tigris River basin.

gel-cheragh.

(None).

54. Gobio gobio (Linnaeus, 1758) - Tedzhen (= Hari) River basin.

mahi kopur kafzi.

(Turkestan gudgeon).

55. Gobio persus Günther, 1899 - Lake Orumiyeh and Caspian Sea basins.

?

(Kura or Persian gudgeon).

56. Hemiculter leucisculus (Basilewski, 1855) - Introduced to the Caspian Sea basin.

tizeh-kuli.

(sharpbelly, common sawbelly, knifefish).

57. Hemigrammocapoeta elegans (Günther, 1868) - Tigris River basin.

?

(None).

58. Hypophthalmichthys molitrix (Valenciennes in Cuvier and Valenciennes, 1844) - Widely introduced.

fytofag (in Khuzestan), kopur noqreai.

(silver carp, thickforehead, white thickforehead).

59. Hypophthalmichthys nobilis (Richardson, 1844) - Introduced to the Caspian Sea basin. sarghundeh (in the Caspian Sea basin), kopur sarkhundeh.

(bighead, painted thickforehead).

60. Iranocypris typhlops Bruun and Kaiser, 1944 - Tigris River basin.

mahi-ye kurghar.

(Iran cave barb, blind cave fish).

61. Leucaspius delineatus (Heckel, 1843) - Caspian Sea basin.

mahi riz noqrehai.

(sunbleak, white aspe, moderlieschen).

62. Pelecus cultratus (Linnaeus, 1758) - Caspian Sea basin.

shamshir mahi, shamshir mahi ab shirin.

(razorfish, sabre carp, sabrefish, ziege).

63. Petroleuciscus persidis (Coad, 1981) - Kor River, Gulf and Hormuz basins.

?

(None).

64. Petroleuciscus ulanus (Günther, 1899) - Lake Orumiyeh basin.

?

(None).

65. Pimephales promelas Rafinesque, 1820 - Introduced to the Namak Lake basin.

?

(fathead minnow).

66. Pseudorasbora parva (Temminck and Schlegel in Siebold, 1842) - Widely introduced.

amurcheh.

(stone moroko; topmouth gudgeon or minnow).

67. Rhodeus amarus (Bloch, 1782) - Caspian Sea basin.

mahi mokhrj lulehai.

(bitterling).

68. Rutilus frisii (Nordmann, 1840) - Caspian Sea basin.

mahi safid, mahi safid daryacheh khazar, sifid mahi (in Gilaki), talaji (in Mazanderani).

(Southern Caspian roach, kutum).

69. Rutilus rutilus (Linnaeus, 1758) - Caspian Sea basin.

kolme, kolme Gorgan, talaji, mahi cheshm qermez.

(roach, vobla; North Caspian, Astrakhan, Kura, Astrabad or Turkmenian vobla or roach).

70. Scardinius erythrophthalmus (Linnaeus, 1758) - Caspian Sea basin.

sorkh pareh, sorkh baleh.

(rudd, redeye, redfin, pearl roach).

71. Schizocypris altidorsalis Bianco and Banarescu, 1982 - Sistan basin.

gorgak.

(None).

72. Schizopygopsis stoliczkai Steindachner, 1866 - Sistan basin.

kopur barfi.

(false osman, Pamir snowcarp).

73. Schizothorax intermedius (McClelland, 1842) - Sistan basin.

mawda (in Sistan), marinka.

(common marinka).

74. Schizothorax pelzami Kessler, 1870 - Tedzhen (= Hari) River and Kavir basin.

khaju, shir mahi.

(Transcaspian marinka).

75. Schizothorax zarudnyi (Nikolskii, 1897) - Sistan basin.

hamun mahi, mahi khaju, shir mahi, anjac.

(None).

76. Squalius cephalus (Linnaeus, 1758) - Caspian Sea, Lake Orumiyeh, Namak Lake and Tigris River basins.

mahi safid (in the Caspian basin and in central Iran); safid rudkhanehai (in Khuzestan).

(Caucasian or European chub).

77. Squalius latus Keyserling, 1861 - Tedzhen (= Hari) River basin.

?

(Transcaspian or Murgab dace).

78. Squalius lepidus Heckel, 1843 - Tigris River basin.

kawar.

(None).

79. Tinca tinca (Linnaeus, 1758) - Caspian Sea basin.

lai mahi, naji, tilkhos.

(tench, green tench).

80. Vimba vimba (Linnaeus, 1758) - Caspian Sea basin.

siah kuli, kuli.

(Caspian vimba, southern white-eye, vimba).

Family 8. Cobitidae (loaches)

1. Cobitis linea (Heckel, 1849) - Kor River, Gulf and Hormuz basins.

sag mahi.

(None).

2. Cobitis taenia Linnaeus, 1758 - Caspian Sea and Tigris River basins.

sag mahi sangi, mahi roshtegar; loch or gel khorok (in Khuzestan).

(spined, spine or spiny loach; stone loach, weatherfish).

3. Sabanejewia aurata (De Filippi, 1865) - Caspian Sea and Tedzhen (= Hari) River basins.

sag mahi khardar.

(Hither Asia spined loach, golden spined or spiny loach).

4. Sabanejewia caspia (Eichwald, 1838) - Caspian Sea basin.

sag mahi khardar, mahi roshtegar talaee.

(Caspian spined loach, spiny loach).

Family 9. Balitoridae (hillstream loaches)

1. Barbatula angorae (Steindachner, 1897) - Caspian Sea, Lake Orumiyeh, Namak Lake and Tigris River basins.

sag mahi Angora, mar mahi.

(Angora loach).

2. Barbatula bergiana (Derzhavin, 1934) - Caspian Sea basin and Namak Lake basin.

sag mahi Safidrud, mar mahi.

(Sefidrud loach).

3. Barbatula brandtii (Kessler, 1877) - Caspian Sea basin.

sag mahi Kura, mar mahi.

(Kura loach).

4. Barbatula farsica (Nalbant and Bianco, 1998) - Namak Lake, Kor River and Gulf basins.

?

(None)

5. Barbatula frenata (Heckel, 1843) - Tigris River basin.

sag mahi, mar mahi.

(None).

6. Barbatula kermanshahensis (Bănărescu and Nalbant, 1967) - Tigris River basin.

sag mahi Kermanshaheh, mar mahi.

(None).

7. Barbatula persa (Heckel, 1849) - Kor River and Lake Maharlu basins.

sag mahi Fars, mar mahi.

(None).

8. Ilamnemacheilus longipinnis Coad and Nalbant, 2005 - Tigris River basin.

?

None.

9. Paracobitis iranica Nalbant and Bianco, 1998 - Namak Lake basin.

?

(None).

10. Paracobitis longicauda (Kessler, 1872) - Tedzhen (= Hari) River basin.

sag mahi kakl dar sharqeh, mar mahi.

(eastern crested loach).

11. Paracobitis malapterura (Valenciennes in Cuvier and Valenciennes, 1846) - Tigris River, Caspian Sea, Tedzhen (= Hari) River, Lake Orumiyeh, Namak Lake and Kor River basins.

sag mahi kakl dar gharbeh, mar mahi.

(western crested loach).

12. Paracobitis rhadinaea (Regan, 1906) - Sistan, Caspian Sea and Tedzhen (= Hari) River basins.

sag mahi, mar mahi.

(None).

13. Paracobitis smithi (Greenwood, 1976) - Tigris River basin.

sag mahi gharzeh, mar mahi.

(blind loach).

14. Paracobitis vignai Nalbant and Bianco, 1998 - Sistan basin

?

(None).

15. Schistura bampurensis (Nikolskii, 1899) - Hamun-e Jaz Murian, Makran and Lut basins.

sag mahi Bampur, mar mahi.

(None).

16. Schistura cristata (Berg, 1898) - Tedzhen (= Hari) River basin and Caspian Sea basins.

sag mahi kakl dartrkmni, mar mahi.

(Turkmenian crested loach).

17. Schistura kessleri (Günther, 1889) - Tedzhen (= Hari) River, Sistan, Mashkid River and Bejestan basins.

sag mahi Bejestan, mar mahi.

(Kessler's loach).

18. Schistura nielseni Nalbant and Bianco, 1998 - Gulf basin.

?

(None)

19. Schistura sargadensis (Nikolskii, 1899) - Hamun-e Jaz Murian, Mashkid River, Bejestan, Kerman-Na'in, Lut and Sirjan basins.

sag mahi Sarhad, mar mahi.

(Turkmen loach).

20. Triplophysa stoliczkai (Steindachner, 1866) - Sistan basin.

sag mahi pamir, mar mahi.

(Tibetan, Pamirs or Stoliczka's loach).

21. Seminemacheilus tongiorgii Nalbant and Bianco, 1998 - Hormuz basin.

?

(None).

Order 8. Silurifomes

Family 10. Bagridae (bagrid catfishes)

1. Mystus pelusius (Solander in Russell, 1794) - Tigris River basin.

abu-zummair, sag mahi, chamu.

(None).

Family 11. Siluridae (sheatfishes)

1. Silurus glanis Linnaeus, 1758 - Caspian Sea, Lake Orumiyeh, Tedzhen (= Hari) River and Tigris River basins.

esbele (in Gilaki), esbele Europaiye, gorba-mahi, gorbeh mahi rogahee, mahi-e sebili; nake (in the Lake Orumiyeh basin).

(European or Danube catfish, sheatfish, wels).

2. Silurus triostegus Heckel, 1843 - Tigris River basin.

esbele, jirri, yary, esbele binonehrin; sag mahi (in Khuzestan).

(Mesopotamian catfish).

Family 12. Sisoridae (sisorid catfishes)

1. Glyptothorax kurdistanicus (Berg, 1931) - Tigris River basin.

gorbeh mahi Kordestan, arteshi.

(None).

2. Glyptothorax silviae Coad, 1981 - Tigris River and Gulf basins.

gorbeh mahi jonub, arteshi.

(None).

Family 13. Heteropneustidae (stinging catfishes)

1. Heteropneustes fossilis (Bloch, 1794) - Introduced to the Tigris River basin.

eshlambo (in Mazandaran); dood or dudeh, abu shalambo (in Khuzestan).

(Indian stinging catfish, stinging catfish).

Order 9. Esociformes

Family 14. Esocidae (pikes)

1. Esox lucius Linnaeus, 1758 - Caspian Sea basin; widely introduced.

ordak mahi, shook chehkhab.

(pike, northern pike).

Order 10. Salmoniformes

Family 15. Salmonidae (salmons and trouts)

1. Coregonus lavaretus (Linnaeus, 1758) - Introduced to the Namak Lake basin.

safid mahi, safid mahi juibarye.

(powan, houting, skelly, gwyniad, lavaret, pollan).

2. Oncorhynchus keta (Walbaum, 1792) - Introduced to the Caspian Sea basin.

azad mahi keta, mahi azad chum.

(chum or dog salmon, keta, summer chum).

3. Oncorhynchus mykiss (Walbaum, 1792) - Widely introduced.

qezel ala ranginkaman; mahi qezel ala, qezel ala (probably in confusion with native Salmo trutta).

(rainbow or steelhead trout).

4. Salmo trutta Linnaeus, 1758 - Caspian Sea, Namak Lake and Lake Orumiyeh basins; widely introduced.

mahi azad or mahi-ye azad, azad mahi (in Gilaki), mahi qezel ala-ye daryacheh khazar (for the subspecies caspius); mahi qezel ala, mahi azad-ye kezel ala-ye bumi, qezel ala khalqermez, qezel ala khalqermez yabumi, qezel ala-ye juibarye (for freshwater residents); mahi azade khal sorkh; qezel ala-ye Liqvanchai (for trout in the Lake Orumiyeh basin).

(Caspian salmon for sea-run fish; brook, brown or river trout).

5. Salvelinus fontinalis (Mitchill, 1814) - Introduced to the Namak Lake basin.

azad mahi cheshmahi.

(brook or speckled trout, brook charr).

6. Stenodus leucichthys (Gueldenstaedt, 1771) - Caspian Sea basin.

safid mahi, azad mahi Volga, mahi ziba, inkonu.

(inconnu, Caspian inconnu, white salmon).

Order 11. Gadiformes

Family 16. Gadidae (cods)

1. Lota lota Linnaeus, 1758 - Caspian Sea basin.

mahi charb, lot.

(burbot, eelpout).

Order 12. Mugiliformes

Family 17. Mugilidae (mullets)

1. Liza abu (Heckel, 1843) - Tigris River and Gulf basins.

biah, biah zury, shochy, or do'kelki in Khuzestan, ? derbak.

(abu mullet).

2. Liza aurata (Risso, 1810) - Introduced to the Caspian Sea basin.

kefal-e auratus or kafal-e auratus, kefal-e zarin, kefal talaee.

(golden or long-finned mullet; golden grey mullet; long-finned grey mullet).

3. Liza saliens (Risso, 1810) - Introduced to the Caspian Sea basin.

kefal-e saliens or kafal-e saliens, kafal-e poozehbank.

(sharpnose, small or grey mullet; leaping grey mullet).

4. Mugil cephalus Linnaeus, 1758 - Tigris River, Gulf, Hormuz and Makran basins.

kefal or kafal, biah srpehn.

(striped, grey, black, jumping, flathead grey, sea or river mullet).

Order 13. Atheriniformes

Family 18. Atherinidae(silversides)

1. Atherina boyeri Risso, 1810 - Caspian Sea basin.

shisheh mahi, gol azin mahi, atrinka.

(Caspian silverside, Caspian sand smelt, Boyer's sand smelt).

Family 19. Cyprinodontidae (tooth-carps)

1. Aphanius dispar (Rüppell, 1829) - Tigris River, Gulf, Hormuz, Makran, Hamun-e Jaz Murian and Mashkid River basins.

gour-e khar, kapurdandan-e balmband.

(high-finned pupfish, Arabian killifish, mother of pearl fish).

2. Aphanius ginaonis (Holly, 1929) - Hormuz basin.

gour-e khar, kapurdandan-e Genu.

(Holly's pupfish).

3. Aphanius isfahanensis Hrbek, Keivany and Coad, 2007

kapurdandan-e Esfahan.

Esfahan tooth-carp).

4. Aphanius mento (Heckel, 1843) - Tigris River basin.

gour-e khar, kapurdandan-e irani.

(Persian pupfish, Persian minnow).

5. Aphanius persicus (Jenkins, 1910) - Lake Maharlu basin.

gour-e khar, kapurdandan-e Fars.

(None).

6. Aphanius sophiae (Heckel, 1849) - Kor River basin.

gour-e khar, kapurdandan-e sufieh.

(Cypris pupfish).

7. Aphanius vladykovi Coad, 1988 - Tigris River basin.

gour-e khar, kapurdandan-e Zagros, mahi parchami.

(Zagros pupfish).

Family 20. Poeciliidae (livebearers)

1. Gambusia holbrooki Girard, 1859 - Widely introduced.

gambusia.

(mosquitofish, eastern mosquitofish).

Order 14. Gasterosteiformes

Family 21. Gasterosteidae (sticklebacks)

1. Gasterosteus aculeatus Linnaeus, 1758 - Introduced to the Caspian Sea basin.

mahi sekhareh.

(threespine stickleback).

2. Pungitius platygaster (Kessler, 1859) - Caspian Sea basin.

nokhar, mahi sehkhareh nama, mahi khardar.

(southern ninespine stickleback).

Family 22. Syngnathidae (pipefishes)

1. Syngnathus abaster Risso, 1826 - Caspian Sea basin.

ney mahi, ney mahi darya-ye Khazar, suzan mahi.

(Caspian or short-snouted pipefish).

Order 15. Synbranchiformes

Family 23. Mastacembelidae (spiny eels)

1. Mastacembelus mastacembelus (Banks and Solander in Russell, 1794) - Tigris River, Kor River and Gulf basins.

mar mahi, mar mahi khardar.

(Mesopotamian spiny eel).

Order 16. Perciformes

Family 24. Centrarchidae (sunfishes)

1. Micropterus salmoides (Lacèpede, 1802) - Introduced to the Tigris River basin.

khorshid mahi baleh kuchak, bs dehanbozorg.

(largemouth bass).

Family 25. Percidae (perches)

1. Perca fluviatilis Linnaeus, 1758 - Caspian Sea basin.

mahi-ye khardar, bacheh suf, mahi suf rudkhaneh-ye Astrakhan, suf Haji Tarkhan, suf rudkhaneh-ye.

(perch; European or Eurasian perch).

2. Sander lucioperca (Linnaeus, 1758) - Caspian Sea basin; widely introduced.

suf, suf-e ma'muli, sibey or sibeyak (in Gilaki), ? sevideh.

(pike-perch, European pikeperch, zander).

3. Sander marinum (Cuvier in Cuvier and Valenciennes, 1828) - Caspian Sea basin.

suf-e daryai-ye.

(sea pike-perch, sea zander, estuarine perch).

Family 26. Sparidae (porgies)

1. Acanthopagrus latus (Houttuyn, 1782) - Gulf basin.

shaanak.

(yellow-finned porgy or seabream; yellow-finned black porgy, Japanese silver bream).

Family 27. Cichlidae (cichlids)

1. Iranocichla hormuzensis Coad, 1982 - Hormuz basin.

siklid Irani, siklid Hormuz.

(None).

Family 28. Gobiidae (gobies)

1. Anatirostrum profundorum (Berg, 1927) - Caspian Sea basin.

gav mahi nokhardkeh.

(duckbill tadpole goby).

2. Benthophilus baeri Kessler, 1877 - Caspian Sea basin.

gav mahi Baer, sag mahi.

(Baer tadpole goby).

3. Benthophilus ctenolepidus Kessler, 1877 - Caspian Sea basin.

gav mahi shfaf, sag mahi.

(transparent tadpole goby).

4. Benthophilus macrocephalus (Pallas, 1787) - Caspian Sea basin.

gav mahi vzgheh, sag mahi.

(Caspian tadpole goby, bighead goby).

5. Benthophilus magistri Iljin, 1927 - Caspian Sea basin.

gav mahi Lenkoran, sag mahi.

(Lenkoran tadpole goby).

6. Benthophilus stellatus (Sauvage, 1874) - Caspian Sea basin.

gav mahi khtri, gav mahi bacheh qurbaqei, sag mahi.

(stellate tadpole goby, Caspian starry goby).

7. Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837 - Tigris River, Gulf, Hormuz and Makran basins.

eshlambo, gel khorok.

(mudskipper).

8. Glossogobius giuris (Hamilton-Buchanan, 1822) - Tigris River, Gulf, Hormuz and Makran basins.

gel-ye mahi, chshm noarye.

(bar-eyed, tank, white or flathead goby).

9. Knipowitschia caucasica (Kavraiskii in Berg, 1916) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi qffazye.

(Caucasus bald goby).

10. Knipowitschia iljini Berg, 1931 - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi.

(None).

11. Neogobius caspius (Eichwald, 1831) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi Khazari.

(Hyrcanian or Caspian goby).

12. Neogobius cyrius (Kessler, 1874) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi.

(None).

13. Neogobius fluviatilis (Pallas, 1814) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi rudkhaneha-ye.

(Caspian sand goby, darkie, Caspian monkey goby).

14. Neogobius gorlap Il'in in Berg, 1949

mahi kafzi, gel-ye mahi sar bozorg, gav mahi sar gondeh.

(bighead goby, gorlap goby).

15. Neogobius melanostomus (Pallas, 1814) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi gerd.

(Caspian round goby, black-spotted goby).

16. Neogobius ratan (Nordmann, 1840) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi ratan.

(Caspian ratan or rotan goby).

17. Periophthalmus waltoni Koumans, 1955 - Tigris River, Gulf, Hormuz and Makran basins, gel khorok, gel-cheragh.

gel-cheragh, mahi-sag.

(mudskipper).

18. Proterorhinus marmoratus (Pallas, 18l4) - Caspian Sea basin.

gav mahi, sag mahi, gel-ye mahi marmari.

(tubenose, marbled or mottled goby).

19. Rhinogobius similis Gill, 1859 - Tedzhen (= Hari) River basin.

gel-ye mahi.

(lake goby).

Family 29. Channidae (snakeheads)

1. Channa gachua (Hamilton-Buchanan, 1822) - Hamun-e Jaz Murian basin.

mahi sar mary.

(brown or smooth-breasted snakehead).

Checklist of Farsi Names

correlate with list above and with species accounts?

abu henesh = Barbus kosswigi

abu shalambo (in Khuzestan) = Heteropneustes fossilis

abu-zummair = Mystus pelusius

alaf khoar = Ctenopharyngodon idella

amur (in Khuzestan) = Ctenopharyngodon idella

amurcheh = Pseudorasbora parva

anjac = Schizothorax zarudnyi

anjarak = Garra rossica

anzeh (at Ahvaz and in southern Khuzestan) = Barbus esocinus

arteshi = Glyptothorax spp.

aslak (in Mazandaran) = Chalcalburnus chalcoides

atrinka = Atherina boyeri

azad mahi (in Gilaki) = Salmo trutta

azad mahi cheshmahi = Salvelinus fontinalis

azad mahi keta = Oncorhynchus keta

azad mahi Volga = Stenodus leucichthys

brk = Tenualosa ilisha

bach (at Ahvaz and in southern Khuzestan) = Barbus esocinus

bacheh suf = Perca fluviatilis

balzard = Barbus esocinus

barg bidy = Capoeta trutta

bas dehanbozorg = Micropterus salmoides

bassan = Barbus pectoralis

beluga = Huso huso

berzem = Barbus barbulus

berzem = Barbus kersin

berzem = Barbus pectoralis

berzem = Capoeta trutta

berzem lab-pahn (in Khuzestan) = Barbus barbulus

biah = Liza abu

biah srpehn = Mugil cephalus

biah zury (in Khuzestan) = Liza abu

binni = Barbus sharpeyi

bizz = Barbus esocinus

blizem = Barbus lacerta

boluga = Huso huso

botak (in Khuzestan) = Cyprinion macrostomum

boz mahi = Barbus barbulus

botak dehan buzorg = Cyprinion macrostomum

butak-e dehan buzorg = Cyprinion macrostomum

butak-e dehan kuchek = Cyprinion kais

butok = Cyprinion kais

chalbash = Acipenser gueldenstaedtii

chamu = Mystus pelusius

darakul = Acipenser persicus

darakul = Acipenser stellatus

derbak = Liza abu

do'kelki (in Khuzestan)= Liza abu

dood (in Khuzestan) = Heteropneustes fossilis

dudeh (in Khuzestan) = Heteropneustes fossilis

esbele (in Gilaki) = Silurus glanis

esbele binonehrin = Silurus triostegus

esbele Europaiye = Silurus glanis

esbele = Silurus triostegus

eshlambo = Boleophthalmus dussumieri

eshlambo = catfishes

eshlambo (in Mazandaran) = Heteropneustes fossilis

eshlambo = Boleophthalmus dussumieri

fil mahi = Huso huso

fytofag (in Khuzestan) = Hypophthalmichthys molitrix

galuk = Cyprinion macrostomum

gambusia = Gambusia holbrooki

gararufa = Garra rufa

gatan = Barbus xanthopterus

gav mahi Baer = Benthophilus baeri

gav mahi khtri = Benthophilus stellatus

gav mahi bacheh qurbaqei = Benthophilus stellatus

gav mahi Lenkoran = Benthophilus magistri

gav mahi nokhardkeh = Anatirostrum profundorum

gav mahi sar gondeh = Neogobius gorlap

gav mahi shfaf = Benthophilus ctenolepidus

gav mahi vzgheh = Benthophilus macrocephalus

gav mahi = Gobiidae

gel-cheragh (in Khuzestan) = Capoeta damascina

gel-cheragh (in the south of Iran) = Capoeta spp.

gel-cheragh = Garra rufa

gel-cheragh = Garra spp.

gel-cheragh = Periophthalmus waltoni

gel khorok = Boleophthalmus dussumieri

gel khorok = Capoeta damascina

gel khorok = Cobitis taenia

gel khorok = Garra rufa

gel khorok = Periophthalmus waltoni

gel-ye mahi = Knipowitschia iljini

gel-ye mahi = Neogobius cyrius

gel-ye mahi = Rhinogobius similis

gel-ye mahi chshm noarye = Glossogobius giuris

gel-ye mahi gerd = Neogobius melanostomus

gel-ye mahi Khazari = Neogobius caspius

gel-ye mahi marmari = Proterorhinus marmoratus

gel-ye mahi qffazye = Knipowitschia caucasica

gel-ye mahi ratan = Neogobius ratan

gel-ye mahi rudkhaneha-ye = Neogobius fluviatilis

gel-ye mahi sar bozorg = Neogobius gorlap

gol azin mahi = Atherina boyeri

gorba-mahi = Silurus glanis

gorbeh mahi jonub = Glyptothorax silviae

gorbeh mahi = Glyptothorax spp.

gorbeh mahi Kordestan = Glyptothorax kurdistanicus

gorbeh mahi rogahee = Silurus glanis

gorgak = Schizocypris altidorsalis

gour-e khar = Aphanius spp.

hamun mahi = Schizothorax zarudnyi

heif-e noon (in Khuzestan) = Chondrostoma regium

himri = Barbus luteus

inkonu = Stenodus leucichthys

jirri = Silurus triostegus

jokhorak = Chondrostoma regium

kafal = Mugil cephalus

kafal-e auratus = Liza aurata

kafal-e poozehbank = Liza saliens

kafal-e saliens = Liza saliens

kafal talaee = Liza aurata

kalashpa = Acanthobrama marmid

kapur (in Gilaki) = Cyprinus carpio

kapurdandan-e Arvand = Aphanius mento

kapurdandan-e balmband = Aphanius dispar

kapurdandan-e Esfahan = Aphanius isfahanensis

kapurdandan-e Fars = Aphanius persicus

kapurdandan-e Genu = Aphanius ginaonis

kapurdandan-e irani = Aphanius mento

kapurdandan-e safiyeh = Aphanius sophiae

kapurdandan-e sufieh = Aphanius sophiae

kapurdandan-e Zagros = Aphanius vladykovi

kara burun = Acipenser persicus

karaz (in Mazandaran) = Carassius auratus

kaviari rusi = Acipenser gueldenstaedtii

kawar = Leuciscus lepidus

kefal = Liza spp.

kefal = Mugil spp.

kefal-e auratus = Liza aurata

kefal-e saliens = Liza saliens

kefal-e zarin = Liza aurata

kefal talaee = Liza aurata

kelka = Clupeonella spp.

keshdi = Acipenser nudiventris

khaju = Schizothorax pelzami

khameh mahi = Chanos chanos

khasham = Aspius aspius

khorshid mahi baleh kuchak = Micropterus salmoides

kilka = Clupeonella spp.

kilka-ye anchovy = Clupeonella engrauliformis

kilka-ye cheshm dorosht = Clupeonella grimmi

kilka-ye ma'muli = Clupeonella cultriventris

kolme = Rutilus rutilus

kolme Gorgan = Rutilus rutilus

kooseh = Carcharhinus leucas

kopur = Cyprinion macrostomum

kopur = Cyprinus carpio

kopur alaf khoar = Ctenopharyngodon idella

kopur barfi = Schizopygopsis stoliczkai

kopur-cheh (in Mazandaran) = Carassius auratus

kopur-e aineheh = Cyprinus carpio

kopur-e rashti (in Khuzestan) = Cyprinus carpio

kopur ma'muli = Cyprinus carpio

kopur noqreai = Hypophthalmichthys molitrix

kopurdandan-e balmband = Aphanius dispar

kapurdandan-e Esfahan = Aphanius isfahanensis

kopurdandan-e Fars = Aphanius persicus

kopurdandan-e Genu = Aphanius ginaonis

kopurdandar Irani = Aphanius mento

kopurdandan-e sufieh = Aphanius sophiae

kopurdandan-e Zagros = Aphanius vladykovi

kopur sarkhundeh = Hypophthalmichthys nobilis

koregon = Coregonus lavaretus

kuli = Vimba vimba

kuli Irani = Alburnus alburnus

kuli Kura = Alburnus filippii

kuli-ye Irani = Alburnus alburnus

kuli-ye Kura = Alburnus filippii

lab-pahn = Barbus barbulus

lai mahi = Tinca tinca

lapak (in Mazandaran) = Alburnoides bipunctatus

loch (in Khuzestan) = Cobitis taenia

lot = Lota lota

lotak = Cyprinion kais

lotak (in Khuzestan) = Cyprinion macrostomum

lotak dehan buzorg = Cyprinion macrostomum

mahi azad (sea run) = Salmo trutta

mahi azad chum = Oncorhynchus keta

mahi azad-ye qezel ala-ye bumi = Salmo trutta

mahi azade khal sorkh = Salmo trutta

mahi charb = Lota lota

mahi cheshm qermez = Rutilus rutilus

mahi dahangerd daryacheh-ye khazar = Caspiomyzon wagneri

mahi-e kaviar = Acipenser gueldenstaedti

mahi-e kaviar = Acipenser nudiventris

mahi-e kaviar = Acipenser persicus

mahi-e kaviar = Acipenser stellatus

mahi-e kaviar = Huso huso

mahi-e sebili = Silurus glanis

mahi howz noqreai = Carassius auratus

mahi kafzi = Neogobius gorlap

mahi kaviar-e bozorg = Huso huso

mahi khaju = Schizothorax zarudnyi

mahi khardar = Pungitius platygaster

mahi khayateh = Alburnoides bipunctatus

mahi khiateh = Alburnoides bipunctatus

mahi khor kuchiku = Tenualosa ilisha

mahi kopur kafzi = Gobio gobio

mahi mokhrj lulehai = Rhodeus amarus

mahi morvarid = Alburnus alburnus

mahi parchami = Aphanius vladykovi

mahi qachaq = bootleg fish

mahi qezel ala = Oncorhynchus mykiss

mahi qezel ala (freshwater resident) = Salmo trutta

mahi qezel ala-ye daryacheh khazar (for the subspecies caspius) = Salmo trutta

mahi riz noqrehai = Leucaspius delineatus

mahi roshtegar = Cobitis taenia

mahi roshtegar talaee = Sabanejewia caspia

mahi safid (Caspian basin and central Iran) = Leuciscus cephalus

mahi safid = Rutilus frisii

mahi safid daryacheh khazar = Rutilus frisii

mahi-sag = Periophthalmus waltoni

mahi sar mary = Channa gachua

mahi sekhareh = Gasterosteus aculeatus

mahi sehkhareh nama = Pungitius platygaster

mahi siah = Barbus mursa

mahi sibeli (at Karaj) = Capoeta buhsei

mahi sim kondpuzh = Abramis sapa

mahi sim nama = Blicca bjoerkna

mahi suf rudkhaneh-y Astrakhan = Perca fluviatilis

mahi-ye azad (sea run) = Salmo trutta

mahi-ye gul = Cyprinus carpio

mahi-ye howz = Carassius auratus

mahi-ye khardar = Perca fluviatilis

mahi-ye kurghar = Iranocypris typhlops

mahi-ye sim = Abramis brama

mahi-ye talaee (in Khuzestan) = Carassius auratus

mahi ziba = Stenodus leucichthys

mar mahi ma'muli = Anguilla anguilla

mar mahi = Caspiomyzon wagneri

mar mahi = Mastacembelus mastacembelus

mar mahi = balitorid spp.

mar mahi dahangerd = Caspiomyzon wagneri

mar mahi khardar = Mastacembelus mastacembelus

marinka = Schizothorax intermedius

mash mahi = Aspius aspius

mawda (in Sistan) = Schizothorax intermedius

naji = Tinca tinca

nake (in the Lake Orumiyeh basin) = Silurus glanis

narbach = Barbus esocinus

nazok = Chondrostoma regium

ney mahi = Syngnathus abaster

ney mahi darya-ye Khazar = Syngnathus abaster

nokhar = Pungitius platygaster

ooshin = Carassius auratus

ordak mahi = Esox lucius

osetra = Acipenser gueldenstaedti

ourange = Barbus spp.

aparak = Alburnoides bipunctatus

posht siah = Alosa pontica

pulad mahi = Barbus capito

puzanok = Alosa caspia

puzanok = Alosa pontica

puze draz = Acipenser stellatus

qezel ala (in Chahar Mahall) = Capoeta damascina

qezel ala = Oncorhynchus mykiss

qezel ala (freshwater resident) = Salmo trutta

qezel ala-ye juibarye (freshwater residents) = Salmo trutta

qezel ala khalqermez (freshwater resident) = Salmo trutta

qezel ala khalqermez (freshwater resident) = Salmo trutta

qezel ala khalqermez yabumi = Salmo trutta

qezel ala Liqvanchai (Lake Orumiyeh basin) = Salmo trutta

qezel ala ranginkaman = Oncorhynchus mykiss

rnj = Barbus capito

rashti (in Khuzestan) = Cyprinus carpio

rizeh keraye = Clupeonella cultriventris

rizeh keraye = Clupeonella engrauliformis

rizeh kuli = Clupeonella cultriventris

rudkhaneh (in Khuzestan) = Leuciscus cephalus

rumi = Barbus grypus

saba = Abramis sapa

safid kuli = Chalcalburnus chalcoides

safid mahi = Coregonus lavaretus

safid mahi = Stenodus leucichthys

safid mahi juibarye = Coregonus lavaretus

safid rudkhanehai (in Khuzestan) = Leuciscus cephalus

sag mahi = Acipenser spp.

sag mahi = Carcharhinus leucas

sag mahi = Cobitis spp.

sag mahi = Huso huso

sag mahi = Gobiidae

sag mahi = Mystus pelusius

sag mahi Angora = Barbatula angorae

sag mahi Bampur = Schistura bampurensis

sag mahi Bejestan = Schistura kessleri

sag mahi Fars = Barbatula persa

sag mahi gharzeh = Paracobitis smithi

sag mahi kakl dar gharbeh = Paracobitis malapterura

sag mahi kakl dar sharqeh = Paracobitis longicauda

sag mahi kakl dartr kmni = Schistura cristata

sag mahi Kermanshaheh = Barbatula kermanshahensis

sag mahi Kura = Barbatula brandtii

sag mahi Pamir = Triplophysa stoliczkai

sag mahi Safidrud = Barbatula bergiana

sag mahi Sarhad = Schistura sargadensis

sag mahi = balitorid spp.

sag mahi (in Khuzestan) = Silurus triostegus

sag mahi khardar = Sabanejewia aurata

sag mahi khardar = Sabanejewia caspia

sag mahi sangi = Cobitis taenia

sangal = Barbus luteus

sarghundeh (in the Caspian basin) = Hypophthalmichthys nobilis

sas mahi = Barbus spp.

sas mahi khazari = Barbus brachycephalus

šep = Acipenser nudiventris

ses mahi = Barbus spp.

ses mahi bozorg = Barbus capito

ses mahi koloft safid rud = Barbus mursa

sevideh = Sander lucioperca

sevroga = Acipenser stellatus

sevruga = Acipenser stellatus

shaanak = Acanthopagrus latus

shaboot = Barbus grypus

shag mahi = Alosa spp.

shag mahi Agrakhan = Alosa sphaerocephala

shag mahi chekameh dar = Alosa caspia

shag mahi-e chesm dorosht = Alosa saposchnikowii

shag mahi darya-ye khazar = Alosa caspia

shag mahi darya-ye siah = Alosa pontica

shag mahi shekam bozorg = Alosa caspia

shag mahi-ye Khazari = Alosa braschnikowii

shag mahi-ye posht siah = Alosa pontica

shah kuli-ye jonubi = Chalcalburnus mossulensis

shah kuli (in Gilaki) = Chalcalburnus chalcoides

shah mahi = Chalcalburnus chalcoides

shamshir mahi = Pelecus cultratus

shamshir mahi ab shirin = Pelecus cultratus

shamshiri = Capoeta buhsei

sheap = Acipenser nudiventris

shebhe-nazy = Acanthobrama marmid

shebhe-sardin = Acanthobrama marmid

shebhe shirbot = Barbus grypus

shebhe shirbot = Barbus kosswigi

shebhe zury (in Khuzestan) = Alburnoides bipunctatus

shelej = Aspius vorax

shenavar = Acipenser nudiventris

shilbot = Barbus grypus

ship = Acipenser nudiventris

shir mahi = Capoeta trutta

shir mahi = Schizothorax pelzami

shir mahi = Schizothorax zarudnyi

shirbot = Barbus grypus

shirbot = Garra rufa

shisheh mahi = Atherina boyeri

shochy (in Khuzestan) = Liza abu

sholge = Aspius vorax

shook chehkhab = Esox lucius

shum = Capoeta aculeata

siah kuli = Chalcalburnus chalcoides

siah kuli = Vimba vimba

siah mahi = Capoeta aculeata

siah mahi Dasht-e Arzhani = Capoeta barroisi

siah mahi = Capoeta capoeta

siah mahi (in the Caspian basin) = Capoeta damascina

siah mahi = Capoeta fusca

siah mahi (in the north of Iran) = Capoeta spp.

sibey (in Gilaki) = Sander lucioperca

sibeyak (in Gilaki) = Sander lucioperca

sifid mahi (in Gilaki) = Rutilus frisii

siklid Hormuz = Iranocichla hormuzensis

siklid Irani = Iranocichla hormuzensis

sim = Abramis brama

sim = Abramis sapa

simparek = Blicca bjoerkna

sobur = Tenualosa ilisha

soleimani = Barbus sharpeyi

soleimani = Barbus subquincunciatus

soong (in northern Khuzestan and Lorestan) = Barbus esocinus

sorkh baleh = Scardinius erythrophthalmus

sorkh pareh = Scardinius erythrophthalmus

sorkheh (in the Zohreh River) = Barbus grypus

soru (in the Dalaki and Shapur river basins) = Capoeta capoeta

sos mahi (in the Dalaki and Shapur river basins) = Barbus barbulus

sos mahi = Barbus spp.

suf = Sander lucioperca

suf-e daryai-ye = Sander marinum

suf-e ma'muli = Sander lucioperca

suf Haji Tarkhan = Perca fluviatilis

suf rudkhaneh-ye = Perca fluviatilis

suzan mahi = Syngnathus abaster

tas mahi = tass mahi

tass mahi = Acipenser spp. (fishing term, includes more than one species)

tass mahi Iran = Acipenser persicus

tass mahi Irani = Acipenser persicus

tass mahi Rus = Acipenser gueldenstaedti

tass mahi shekam brahne = Acipenser nudiventris

talaji (in Mazandarani) = Rutilus frisii

talaji = Rutilus rutilus

tilkhos = Capoeta capoeta

tilkhos = Tinca tinca

tirij = Acipenser persicus

tizeh-kuli = Hemiculter leucisculus

touyeni = Barbus pectoralis

touyeni (in Khuzestan)= Capoeta damascina

touyeni (in the south of Iran) = Capoeta spp.

touyeni (in Khuzestan) = Capoeta trutta

usach bulatmai = Barbus capito

usach chanari = Barbus capito

uzun burun = Acipenser stellatus

yary = Silurus triostegus

zabur = Tenualosa ilisha

zalun (in Gilaki) = Alosa caspia

zalun (in Gilaki) = Alosa pontica

zard pareh = Barbus capito

zardeh = Capoeta damascina

zardehpar = Barbus brachycephalus

zardehpar = Barbus capito

zardek = Barbus brachycephalus

zardek = Barbus capito

zardek = Barbus spp.

zardek-e qalami = Barbus mursa

zabur = Tenualosa ilisha

zomur = Tenualosa ilisha

References

See the Bibliography for a complete listing of works on Iranian fishes. The References here are meant as a source for names and an update of Coad (1995).

Al-Nasiri, S. K. and Hoda, S. M. S. 1976. A guide to the freshwater fishes of Iraq. Basrah Natural History Museum Publication, 1: xii + 124 pp.

Berg, L. S. 1948-1949. Freshwater fishes of the USSR and adjacent countries. Israel Program for Scientific Translations, Jerusalem (1962-1965). 3 volumes.

Coad, B. W. 1995. Freshwater Fishes of Iran. Acta Scientiarum Naturalium Academiae Scientiarum Bohemicae, Brno, 29(1):1-64.

Coad, B. W. 1996a. Systematics of the tooth-carp genus Aphanius Nardo, 1827 (Actinopterygii: Cyprinodontidae) in Fars Province, southern Iran. Biologia, Bratislava, 51(2):163-172.

Coad, B. W. 1996b. Systematics of the shah mahi, Chalcalburnus chalcoides (Güldenstädt, 1772), in the southern Caspian Sea basin (Actinopterygii: Cyprinidae). Zoology in the Middle East, 12:65-70.

Coad, B. W. 1999. Bibliography on the Freshwater Fishes of Iran. First posted 10 September 1996. 190 k. At www.geocities.com/CapeCanaveral/Hall/1345/. Page maintained by Yazdan Keivany, University of Alberta, Edmonton (ykeivany@gpu.srv.ualberta.ca).

Coad, B. W. and Al-Hassan, L. A. J. 1989. A Bibliography of the Fishes of the Tigris-Euphrates Basin / Bibliographie der Fische des Euphrat-Tigris-Beckens. Max Kasparek Verlag, Heidelberg. 56 pp.

Coad, B. W. und Kuru, M. 1986. Bibliographie der Fische der Türkei/A Bibliography of the Fishes of Turkey, pp. 15-77. In: Kasparek, M. (Ed.). Zoologische Bibliographie der Türkei. Zoological Bibliography of Turkey. Pisces, Amphibia, Reptilia. Max Kasparek Verlag, Heidelberg. 118 pp.

Coad, B. W. and Papahn, F. 1988. Shark attacks in the rivers of southern Iran. Environmental Biology of Fishes, 23(1-2):131-134. (Reprinted with same pagination in McAllister, D. E. and Kott, E. (Eds.). On lampreys and fishes: a memorial anthology in honor of Vadim D. Vladykov. Developments in Environmental Biology of Fishes, 8:1-162, Kluwer Academic Publishers, Dordrecht/Boston/London).

Elvira, B. 1997. Taxonomy of the genus Chondrostoma (Osteichthyes, Cyprinidae): an updated review. Folia Zoologica, 46(Supplement 1):1-14.

Eschmeyer, W. N. (Ed.). 1998. Catalog of Fishes. California Academy of Sciences, San Francisco. 3 volumes, CD-ROM.

Heckel, J. J. 1843. Ichthyologie. In: Russegger, J. Reisen in Europa, Asien und Afrika. Stuttgart, 1(2):991-1099, Taf. II-XIII.

Heckel, J. J. 1846-1849. Naturhistorischer Anhang. In: Russegger, J. Reisen in Europa, Asien und Afrika, Stuttgart, 2(3):207-254, Taf. XIV-XXI.

Khalaf, K. T. 1961. The marine and freshwater fishes of Iraq. Ar-Rabitta Press, Baghdad. 164 pp.

Kottelat, M. 1997. European freshwater fishes. An heuristic checklist of the freshwater fishes of Europe (exclusive of former USSR), with an introduction for non-systematists and comments on nomenclature and conservation. Biologia, Bratislava, 52 (Supplement 5):1-271.

Lazara, K. J. 1995. History of the genera Lebia Oken 1817 and Lebias Goldfuss 1820 (Teleostei: Cyprinodontiformes: Cyprinodontidae) with designation of a type species for Lebias. Copeia, 1995(2):501-503.

Lindberg, G. U. and Heard, A. S. 1972. Slovar nazvanii presnovodnikh ryb SSSR na yazikakh narodov SSSR i evropeiskikh stran [Dictionary of the Names of Freshwater Fishes of the USSR on the Languages of the Peoples of the USSR and European Countries]. Izdatel'stvo Nauka, Leningrad. 367 pp.

Mahdi, N. 1962. Fishes of Iraq. Ministry of Education, Baghdad. 82 pp.

Mirza, M. R. 1972. Freshwater fishes of Baluchistan Province, Pakistan. Biologia, Lahore, 18:153-190.

Mirza, M. R. 1978. History of ichthyology in Pakistan. Biologia, Lahore, 24(2):305-348.

Mirza, M. R. 1990. Pakistan main taza pani ki macchalian [Freshwater Fishes of Pakistan]. Urdu Science Board, Lahore. 128 pp.

Nalbant, T. T. and Bianco, P. G. 1998. The loaches of Iran and adjacent regions with description of six new species (Cobitoidea). Italian Journal of Zoology, 65 (Supplement):109-123.

Nelson J. S. 1994. Fishes of the World. 3rd Edition. John Wiley & Sons, New York. xvii + 600 pp.

OECD (Organisation for Economic Co-operation and Development). 1995. Multilingual Dictionary of Fish and Fish Products. Fourth Edition. Fishing News Books, Oxford. xv + 352 pp.

Ricker, W. E. 1973. Russian-English Dictionary for Students of Fisheries and Aquatic Biology. Bulletin of the Fisheries Research Board of Canada, 183:xi + 428 pp.

Robins, C. R., Bailey, R. M., Bond, C. E., Brooker, J. R., Lachner, E. A., Lea, R. N. and Scott, W. B. 1991a. Common and Scientific Names of Fishes from the United States and Canada. Fifth Edition. American Fisheries Society Special Publication, 20:183 pp.

Robins, C. R., Bailey, R. M., Bond, C. E., Brooker, J. R., Lachner, E. A., Lea, R. N. and Scott, W. B. 1991b. World Fishes Important to North Americans Exclusive of Species from the Continental Waters of the United States and Canada. American Fisheries Society Special Publication, 21:viii + 243 pp.


Scientific Names

The first Checklist is for all Iranian species followed by Checklists according to drainage basins. The first and main Checklist gives the classification of the Iranian freshwater fish species following the arrangement of families in Nelson (1994). The Species Accounts do not give a higher classification and are arranged by families and genera.

E = Exotic, M = Marine, entering freshwater from the Persian Gulf, but not normally resident (Caspian Sea species regarded as "freshwater"), N = reported from neighbouring countries, ? = probably a misidentification. Details on sources for these records are in the Species Accounts.

Checklist of Iranian Freshwater Fishes

Class 1. Cephalaspidomorphi

Order 1. Petromyzontiformes

Family 1. Petromyzontidae

Caspiomyzon wagneri (Kessler, 1870)

Class 2. Chondrichthyes

Order 2. Carcharhiniformes

Family 2. Carcharhinidae

M Carcharhinus dussumieri (Valenciennes in Müller and Henle, 1839)

Carcharhinus leucas (Valenciennes in Müller and Henle, 1839)

Class 3. Actinopterygii

Order 3. Acipenseriformes

Family 3. Acipenseridae

E Acipenser baeri Brandt, 1869

Acipenser gueldenstaedtii Brandt in Brandt and Ratzeburg, 1833

Acipenser nudiventris Lovetzky, 1828

Acipenser persicus Borodin, 1897

N Acipenser ruthenus Linnaeus, 1758

Acipenser stellatus Pallas, 1771

Huso huso (Linnaeus, 1758)

N Pseudoscaphirhynchus kaufmanni (Kessler, 1877)

Order 4. Anguilliformes

Family 4. Anguillidae

E Anguilla anguilla (Linnaeus, 1758)

Order 5. Clupeiformes

Family 5. Engraulidae

M Engraulis vitirostris ?

E Engraulis encrasicholus (Linnaeus, 1758)

M Thryssa hamiltoni (Gray, 1835)

M, N Thryssa purava Hamilton-Buchanan, 1822)

M, N Thryssa setirostris (Broussonet, 1782)

M Thryssa whiteheadi Wongratana, 1983

Family 6. Chirocentridae

M, N Chirocentrus nudus Swainson, 1839

Family 7. Clupeidae

Alosa braschnikowii (Borodin, 1904)

Alosa caspia (Eichwald, 1838)

N Alosa curensis (Suvorov, 1907)

Alosa pontica (Eichwald, 1838)

Alosa saposchnikowii (Grimm, 1887)

Alosa sphaerocephala (Berg, 1913)

Clupeonella cultriventris (Nordmann, 1840)

Clupeonella engrauliformis (Borodin, 1904 (?? 1906 in Eschmeyer et al 1996)

Clupeonella grimmi Kessler, 1877

M, N Hilsa kelee (Cuvier, 1829)

M, N Ilisha compressa Randall, 1995

M, N Ilisha megaloptera (Swainson, 1839)

M, N Nematalosa nasus (Bloch, 1795)

Tenualosa ilisha (Hamilton-Buchanan, 1822)

Order 6. Gonorhynchiformes

Family 8. Chanidae

M Chanos chanos (Forsskål, 1775)

Order 7. Cypriniformes

Family 9. Cyprinidae

Abramis brama (Linnaeus, 1758)

Abramis sapa (Pallas, 1814)

Acanthalburnus microlepis (De Filippi, 1863)

Acanthalburnus urmianus (Günther, 1899)

Acanthobrama marmid Heckel, 1843

Alburnoides bipunctatus (Bloch, 1782)

N Alburnoides taeniatus (Kessler, 1874)

Alburnus alburnus (Linnaeus, 1758)

N Alburnus caeruleus Heckel, 1843

Alburnus filippi Kessler, 1877

Aspidoparia morar (Hamilton-Buchanan, 1822)

N Aspiolucius esocinus ?

Aspius aspius (Linnaeus, 1758)

Aspius vorax Heckel, 1843

Barbus barbulus Heckel, 1849

Barbus brachycephalus Kessler, 1872

Barbus capito (Güldenstaedt, 1773)

Barbus esocinus (Heckel, 1843)

Barbus grypus Heckel, 1843

Barbus kersin Heckel, 1843

Barbus kosswigi (Ladiges, 1960)

Barbus lacerta Heckel, 1843

Barbus luteus (Heckel, 1843)

Barbus mursa (Güldenstaedt, 1773)

Barbus pectoralis Heckel, 1843

Barbus sharpeyi Günther, 1874

Barbus sublimus Coad and Najafpour, 1997

Barbus subquincunciatus Günther, 1868

Barbus xanthopterus (Heckel, 1843)

Barilius mesopotamicus Berg, 1932

Blicca bjoerkna (Linnaeus, 1758)

Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

Capoeta barroisi Lortet in Barrois, 1894

Capoeta buhsei Kessler, 1877

Capoeta capoeta (Güldenstaedt, 1773)

Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

Capoeta fusca Nikol'skii, 1897

Capoeta trutta (Heckel, 1843)

N Capoetobrama kuschakewitschi (Kessler, 1872)

E Carassius auratus (Linnaeus, 1758)

E Carassius carassius (Linnaeus, 1758)

Chalcalburnus atropatenae (Berg, 1925)

Chalcalburnus chalcoides (Güldenstaedt, 1772)

Chalcalburnus mossulensis (Heckel, 1843)

Chondrostoma cyri Kessler, 1877

Chondrostoma regium (Heckel, 1843)

Crossocheilus latius (Hamilton-Buchanan, 1822)

E Ctenopharyngodon idella (Valenciennes in Cuvier and Valenciennes, 1844)

Cyprinion kais Heckel, 1843

Cyprinion macrostomum Heckel, 1843

Cyprinion milesi (Day, 1880)

Cyprinion tenuiradius Heckel, 1849

Cyprinion watsoni (Day, 1872)

Cyprinus carpio Linnaeus, 1758

Garra persica Berg, 1913

Garra rossica (Nikol'skii, 1900)

Garra rufa (Heckel, 1843)

Garra variabilis (Heckel, 1843)

Gobio gobio (Linnaeus, 1758)

Gobio persus Günther, 1899

E Hemiculter leucisculus (Basilewsky, 1855)

Hemigrammocapoeta elegans (Günther, 1868)

E Hypophthalmichthys molitrix (Valenciennes in Cuvier and Valenciennes, 1844)

E Hypophthalmichthys nobilis (Richardson, 1844)

Iranocypris typhlops Bruun and Kaiser, 1944

N Labeo dero (Hamilton-Buchanan, 1822)

N Labeo gedrosicus Zugmayer, 1912

N Labeo macmahoni Zugmayer, 1912

Leucaspius delineatus (Heckel, 1843)

Leuciscus cephalus (Linnaeus, 1758)

N Leuciscus idus (Linnaeus, 1758)

Leuciscus latus (Keyserling, 1861)

Leuciscus lepidus (Heckel, 1843)

Petroleuciscus persidis (Coad, 1981)

Petroleuciscus ulanus (Günther, 1899)

E, N Mylopharyngodon piceus (Richardson, 1846)

E, N Parabramis pekinensis (Basilewsky, 1855)

Pelecus cultratus (Linnaeus, 1758)

E Pimephales promelas Rafinesque, 1820

E, N Pseudogobio rivularis (Basilewsky, 1855)

E Pseudorasbora parva (Temminck and Schlegel in Siebold, 1842)

E, N Rhodeus ocellatus (Kner, 1866)

Rhodeus sericeus (Pallas, 1776)

Rutilus frisii (Nordmann, 1840)

Rutilus rutilus (Linnaeus, 1758)

Scardinius erythrophthalmus (Linnaeus, 1758)

Schizocypris altidorsalis Bianco and Banarescu, 1982

Schizopygopsis stoliczkai Steindachner, 1866

N Schizothorax esocinus (Heckel, 1838)

Schizothorax intermedius McClelland, 1842

N Schizothorax labiatus (McClelland, 1842)

Schizothorax pelzami Kessler, 1870

N Schizothorax plagiostomus Heckel, 1838

Schizothorax zarudnyi (Nikol'skii, 1897)

Tinca tinca (Linnaeus, 1758)

Vimba vimba (Linnaeus, 1758)

Family 10. Cobitidae

Cobitis aurata De Filippi, 1863

Cobitis caspia Eichwald, 1838

Cobitis linea (Heckel, 1849)

Cobitis taenia Linnaeus, 1758

N Misgurnus anguillicaudatus (Cantor, 1842)

Family 11. Balitoridae

Barbatula angorae (Steindachner, 1897)

Barbatula bergiana (Derzhavin, 1934)

Barbatula brandti (Kessler, 1877)

Barbatula farsica (Nalbant and Bianco, 1998)

Barbatula frenata (Heckel, 1843)

Barbatula kermanshahensis (Bănărescu and Nalbant, 1967)

Barbatula persa (Heckel, 1849)

Ilamnemacheilus longipinnis Coad and Nalbant, 2005

N Paracobitis boutanensis (McClelland, 1842)

N Paracobitis ghazniensis (Bănărescu and Nalbant, 1967)

Paracobitis iranica Nalbant and Bianco, 1998

Paracobitis longicauda (Kessler, 1872)

Paracobitis malapterura (Valenciennes in Cuvier and Valenciennes, 1846)

Paracobitis rhadinaea (Regan, 1906)

Paracobitis smithi (Greenwood, 1976)

Paracobitis vignai Nalbant and Bianco, 1998

N Schistura baluchiorum (Zugmayer, 1912)

Schistura bampurensis (Nikol'skii, 1899)

Schistura cristata (Berg, 1898)

Schistura kessleri (Günther, 1889)

N Schistura lindbergi (Banarescu and Mirza, 1965)

Schistura nielseni Nalbant and Bianco, 1998

N Schistura prashari (Hora, 1933)

Schistura sargadensis (Nikol'skii, 1899)

Seminemacheilus tongiorgii Nalbant and Bianco, 1998

N Triplophysa farwelli (Hora, 1935)

N Triplophysa griffithi (Günther, 1868)

Triplophysa stoliczkai (Steindachner, 1866)

N Turcinoemacheilus kosswigi Banarescu and Nalbant, 1964

Order 8. Silurifomes

Family 12. Bagridae

Mystus pelusius (Solander in Russell, 1794)

Family 13. Siluridae

Silurus glanis Linnaeus, 1758

Silurus triostegus Heckel, 1843

Family 14. Sisoridae

N Glyptothorax armeniacus (Berg, 1918)

N Glyptothorax cous (Linnaeus, 1766)

Glyptothorax kurdistanicus (Berg, 1931)

Glyptothorax silviae Coad, 1981

N Glyptothorax steindachneri (Pietschmann, 1913)

Family 15. Heteropneustidae

E Heteropneustes fossilis (Bloch, 1794)

Family 16. Ariidae

M Arius thalassinus (Rüppell, 1837)

Family 17. Plotosidae

M Plotosus lineatus (Thunberg, 1787)

Order 9. Esociformes

Family 18. Esocidae

Esox lucius Linnaeus, 1758

Order 10. Salmoniformes

Family 19. Salmonidae

E Coregonus lavaretus (Linnaeus, 1758)

E, N Oncorhynchus gorbuscha (Walbaum, 1792)

E Oncorhynchus keta (Walbaum, 1792)

E, N Oncorhynchus kisutch (Walbaum, 1792)

E Oncorhynchus mykiss (Walbaum, 1792)

Salmo trutta Linnaeus, 1758

E Salvelinus fontinalis (Mitchill, 1814)

Stenodus leucichthys (Güldenstaedt, 1772)

Order 11. Gadiformes

Family 20. Gadidae

Lota lota (Linnaeus, 1758)

Order 12. Mugiliformes

Family 21. Mugilidae

Liza abu (Heckel, 1843)

E Liza aurata (Risso, 1810)

M, N Liza carinata (Valenciennes in Cuvier and Valenciennes, 1836)

M Liza macrolepis (Smith, 1849)

M, N Liza oligolepis (Bleeker, 1859)

E Liza saliens (Risso, 1810)

M, N Liza subviridis (Valenciennes in Cuvier and Valenciennes, 1836)

M, N Liza vaigiensis (Quoy and Gaimard, 1824)

M Mugil cephalus Linnaeus, 1758

Order 13. Atheriniformes

Family 22. Atherinidae

Atherina boyeri Risso, 1810

Order 14. Beloniformes

Family 23. Adrianichthyidae

E, N Oryzias latipes (Temminck and Schlegel, 1846)

Family 24. Belonidae

Strongylura strongylurus (van Hasselt, 1823)

Family 25. Hemiramphidae

M, N Hemiramphus spp.

M, N Hyporhamphus spp.

M, N Rhynchorhamphus georgii (Valenciennes in Cuvier and Valenciennes, 1847)

Order 15. Cyprinodontiformes

Family 26. Poeciliidae

E, N Gambusia affinis (Baird and Girard, 1853)

E Gambusia holbrooki Girard, 1859

Family 27. Cyprinodontidae

Aphanius dispar (Rüppell, 1829)

Aphanius ginaonis (Holly, 1929)

Aphanius mento (Heckel, 1843)

Aphanius persicus (Jenkins, 1910)

Apahnius sophiae (Heckel, 1849)

Aphanius vladykovi Coad, 1988

Order 16. Gasterosteiformes

Family 28. Gasterosteidae

E Gasterosteus aculeatus Linnaeus, 1758

Pungitius platygaster (Kessler, 1859)

N Pungitius platygaster aralensis (Kessler, 1877)

Family 29. Syngnathidae

Syngnathus abaster Risso, 1826

Order 17. Synbranchiformes

Family 30. Mastacembelidae

Mastacembelus mastacembelus (Banks and Solander in Russell, 1794)

Order 18. Scorpaeniformes

Family 31. Platycephalidae

M Platycephalus indicus (Linnaeus, 1758)

Order 19. Perciformes

Family 32. Percichthyidae

E, N Morone saxatilis (Walbaum, 1792)

Family 33. Serranidae

M, N Epinephelus tauvina (Forsskål, 1775)

Family 34. Centrarchidae

E Lepomis macrochirus Rafinesque, 1819

E Micropterus salmoides (Lacèpede, 1802)

Family 35. Percidae

Perca fluviatilis Linnaeus, 1758

Stizostedion lucioperca (Linnaeus, 1758)

Stizostedion marinum (Cuvier in Cuvier and Valenciennes, 1828)

Family 36. Apogonidae

M, N Apogon nigripinnis Cuvier in Cuvier and Valenciennes, 1828

Family 37. Sillaginidae

M Sillago sihama (Forsskål, 1775)

Family 38. Carangidae

M, N Alepes djedaba (Forsskål, 1775)

M, N Caranx sexfasciatus Quoy and Gaimard, 1824

Family 39. Leiognathidae

M, N Leiognathus fasciatus (Lacepède, 1803)

Family 40. Haemulidae

M, N Pomadasys argyreus (Valenciennes in Cuvier and Valenciennes, 1833)

Family 41. Sparidae

M, N Acanthopagrus berda (Forsskål, 1775)

Acanthopagrus latus (Houttuyn, 1782)

Family 42. Polynemidae

M, N Eleutheronema tetradactylum (Shaw, 1804)

Family 43. Sciaenidae

M, N Argyrosomus amoyensis (Bleeker, 1863)

M, N, ? Johnieops aneus (Bloch, 1793)

M, ? Johnius belangerii (Cuvier in Cuvier and Valenciennes, 1830)

M, N, ? Johnius carutta Bloch, 1793

M, N Otolithes ruber (Schneider in Bloch and Schneider, 1801)

Family 44. Mullidae

E, N Mullus barbatus Linnaeus, 1758

Family 45. Terapontidae

M, N Terapon theraps (Cuvier in Cuvier and Valenciennes, 1829)

Family 46. Cichlidae

Iranocichla hormuzensis Coad, 1982

E, N Oreochromis niloticus (Linnaeus, 1758)

E, N Tilapia zilli (Gervais, 1848)

Family 47. Gobiidae

M, N Acentrogobius dayi Koumans, 1955

Anatirostrum profundorum (berg, 1927)

N Asra turcomana Il'in, 1941

M, N Bathygobius fuscus (Rüppell, 1830)

N Benthophiloides brauneri Beling and Il'in, 1927

Benthophilus baeri Kessler, 1877

Benthophilus ctenolepiDus Kessler, 1877

N Benthophilus granulosus Kessler, 1877

N Benthophilus grimmi Kessler, 1877

N Benthophilus leptocephalus Kessler, 1877

N Benthophilus leptorhynchus Kessler, 1877

Benthophilus macrocephalus (Pallas, 1788)

Benthophilus magistri Il'in, 1927

N Benthophilus mahmudbejovi Ragimov, 1976

N Benthophilus spinosus Kessler, 1877

Benthophilus stellatus (Sauvage, 1874)

N Benthophilus svetovidovi Pinchuk and Ragimov, 1979

Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837

N Caspiosoma caspium (Kessler, 1877)

Glossogobius giuris (Hamilton-Buchanan, 1822)

N Hyrcanogobius bergi Il'in, 1928

Knipowitschia caucasica (Berg, 1916)

Knipowitschia iljini Berg, 1931

N Knipowitschia longecaudata (Kessler, 1877)

N Mesogobius nigronotatus (Kessler, 1877)

N Mesogobius nonultimus (Il'in, 1936)

N Neogobius bathybius (Kessler, 1877)

Neogobius caspius (Eichwald, 1831)

Neogobius cyrius (Kessler, 1874)

Neogobius fluviatilis (Pallas, 1814)

N Neogobius gymnotrachelus (Kessler, 1857)

Neogobius iljini Vasil'eva and Vasil'ev, 1996

Neogobius melanostomus (Pallas, 1814)

Neogobius ratan (Nordmann, 1840)

N Neogobius syrman (Nordmann, 1840)

Periophthalmus waltoni Koumans, 1955

Proterorhinus marmoratus (Pallas, 18l4)

E, N Rhinogobius sp. ? similis Gill, 1859

Scartelaos tenuis (Day, 1876)

Family 48. Ephippididae

M, N Drepane punctata (Linnaeus, 1758)

Family 49. Scatophagidae

Scatophagus argus (Bloch, 1788)

Family 50. Stromateidae

M, N Pampus argenteus (Euphrasen, 1788)

M, N Pampus chinensis (Euphrasen, 1788)

Family 51. Channidae

E, N Channa argus ((Cantor, 1842)

Channa gachua (Hamilton-Buchanan, 1822)

Order 20. Pleuronectiformes

Family 52. Scophthalmidae

E, N Psetta maxima (Linnaeus, 1758)

Family 53. Pleuronectidae

E Platichthys flesus (Linnaeus, 1758)

Family 54. Soleidae

M, N Euryglossa orientalis (Schneider in Bloch and Schneider, 1801)

M, N, ? Solea heinii Steindachner, 1907

Family 55. Cynoglossidae

M, N Cynoglossus arel (Schneider in Bloch and Schneider, 1801)

M, N Cynoglossus lingua (Hamilton-Buchanan, 1822)

Checklists by Drainage Basin for Native Fishes

Lake Orumiyeh

Family Cyprinidae

1. Acanthalburnus urmianus (Günther, 1899)

2. Alburnoides bipunctatus (Bloch, 1782)

3. Barbus lacerta Heckel, 1843

4. Capoeta capoeta (Gueldenstaedt, 1773)

5. Chalcalburnus atropatenae (Berg, 1925)

6. Gobio persus Günther, 1899

7. Leuciscus cephalus (Linnaeus, 1758

8. Leuciscus ulanus Günther, 1899

Family Balitoridae

1. Nemacheilus angorae Steindachner, 1897

2. Nemacheilus malapterurus (Valenciennes in Cuvier and Valenciennes, 1846)

Family Siluridae

1. Silurus glanis Linnaeus, 1758

Family Salmonidae

1. Salmo trutta Linnaeus, 1758

Caspian Sea

Family Petromyzontidae

1. Caspiomyzon wagneri (Kessler, 1870)

Family Acipenseridae

1. Acipenser gueldenstaedtii Brandt, 1833

2. Acipenser nudiventris Lovetzky, 1828

3. Acipenser persicus Borodin, 1897

4. Acipenser stellatus Pallas, 1771

5. Huso huso (Linnaeus, 1758)

Family Anguillidae

1. Anguilla anguilla (Linnaeus, 1758)

Family Clupeidae

1. Alosa braschnikowii (Borodin, 1904)

2. Alosa caspia (Eichwald, 1838)

3. Alosa pontica (Eichwald, 1838)

4. Alosa saposchnikowii (Grimm, 1887)

5. Alosa sphaerocephala (Berg, 1913)

6. Clupeonella cultriventris (Nordmann, 1840)

7. Clupeonella engrauliformis (Borodin, 1904)

8. Clupeonella grimmi Kessler, 1877

Family Cyprinidae

1. Abramis brama (Linnaeus, 1758)

2. Abramis sapa (Pallas, 1814)

3. Acanthalburnus microlepis (De Filippi, 1863)

4. Alburnoides bipunctatus (Bloch, 1782)

5. Alburnus alburnus (Linnaeus, 1758)

6. Alburnus filippi Kessler, 1877

7. Aspius aspius (Linnaeus, 1758)

8. Barbus brachycephalus Kessler, 1872

9. Barbus capito (Gueldenstaedt, 1773)

10. Barbus lacerta Heckel, 1843

11. Barbus mursa (Gueldenstaedt, 1773)

12. Blicca bjoerkna (Linnaeus, 1758)

13. Capoeta capoeta (Gueldenstaedt, 1773)

14. Chalcalburnus chalcoides (Gueldenstaedt, 1772)

15. Chondrostoma cyri Kessler, 1877

16. Cyprinus carpio Linnaeus, 1758

17. Gobio gobio (Linnaeus, 1758)

18. Leucaspius delineatus (Heckel, 1843)

19. Leuciscus cephalus (Linnaeus, 1758

20. Pelecus cultratus (Linnaeus, 1758)

21. Rhodeus sericeus (Pallas, 1776)

22. Rutilus frisii (Nordmann, 1840)

23. Rutilus rutilus (Linnaeus, 1758)

24. Scardinius erythrophthalmus (Linnaeus, 1758)

25. Tinca tinca (Linnaeus, 1758)

26. Vimba vimba (Linnaeus, 1758)

Family Cobitidae

1. Cobitis aurata (De Filippi, 1865)

2. Cobitis caspia Eichwald, 1838

3. Cobitis taenia Linnaeus, 1758

Family Balitoridae

1. Nemacheilus angorae Steindachner, 1897

2. Nemacheilus bergianus Derzhavin, 1934

3. Nemacheilus brandti Kessler, 1877

4. Nemacheilus malapterurus (Valenciennes in Cuvier and Valenciennes, 1846)

Family Siluridae

1. Silurus glanis Linnaeus, 1758

Family Esocidae

1. Esox lucius Linnaeus, 1758

Family Salmonidae

1. Salmo trutta Linnaeus, 1758

2. Stenodus leucichthys (Gueldenstaedt, 1772)

Family Gadidae

1. Lota lota Linnaeus, 1758

Family Atherinidae

1. Atherina boyeri Risso, 1810

Family Gasterosteidae

1. Pungitius platygaster (Kessler, 1859)

Family Syngnathidae

1. Syngnathus abaster Risso, 1826

Family Percidae

1. Perca fluviatilis Linnaeus, 1758

2. Stizostedion lucioperca (Linnaeus, 1758)

3. Stizostedion marinum (Cuvier in Cuvier and Valenciennes, 1828)

Family Gobiidae

1. Benthophilus ctenolepidus Kessler, 1877

2. Benthophilus macrocephalus (Pallas, 1787)

3. Benthophilus magistri Iljin, 1927

4. Benthophilus stellatus (Sauvage, 1874)

5. Knipowitschia caucasica (Kavraiskii in Radde, 1899)

6. Neogobius caspius (Eichwald, 1831)

7. Neogobius fluviatilis (Pallas, 1814)

8. Neogobius iljini Vasil'eva and Vasil'ev, 1996

9. Neogobius melanostomus (Pallas, 1814)

10. Neogobius ratan (Nordmann, 1840)

11. Proterorhinus marmoratus (Pallas, 1814)

Hari River

Family Cyprinidae

1. Alburnoides bipunctatus (Bloch, 1782)

2. Capoeta capoeta (Gueldenstaedt, 1773)

3. Capoeta fusca Nikol'skii, 1897

4. Cyprinus carpio Linnaeus, 1758

5. Garra rossica (Nikol'skii, 1900)

6. Gobio gobio (Linnaeus, 1758)

7. Leuciscus latus (Keyserling, 1861)

8. Schizothorax pelzami Kessler, 1870

Family Cobitidae

1. Cobitis aurata (De Filippi, 1865)

Family Balitoridae

1. Nemacheilus kessleri Günther, 1889

2. Nemacheilus longicauda (Kessler, 1872)

Family Siluridae

1. Silurus glanis Linnaeus, 1758

Tigris River

Family Carcharhinidae

1. Carcharhinus leucas (Valenciennes in MuLler and Henle, 1839)

Family Clupeidae

1. Tenualosa ilisha (Hamilton-Buchanan, 1822)

Family Cyprinidae

1. Acanthobrama marmid Heckel, 1843

2. Alburnoides bipunctatus (Bloch, 1782)

3. Aspius vorax Heckel, 1843

4. Barbus barbulus Heckel, 1849

5. Barbus esocinus (Heckel, 1843)

6. Barbus grypus Heckel, 1843

7. Barbus kersin Heckel, 1843

8. Barbus kosswigi (Ladiges, 1960)

9. Barbus lacerta Heckel, 1843

10. Barbus luteus (Heckel, 1843)

11. Barbus mystaceus (Pallas, 1814)

12. Barbus pectoralis Heckel, 1843

13. Barbus plebejus Bonaparte, 1832

14. Barbus sharpeyi (Günther, 1874)

15. Barbus subquincunciatus Günther, 1868

16. Barbus xanthopterus (Heckel, 1843)

17. Barilius mesopotamicus Berg, 1932

18. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

19. Capoeta barroisi (Lortet in Barrois, 1894)

20. Capoeta capoeta (Gueldenstaedt, 1773)

21. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

22. Capoeta trutta (Heckel, 1843)

23. Chalcalburnus mossulensis (Heckel, 1843)

24. Chondrostoma regium (Heckel, 1843)

25. Cyprinion kais Heckel, 1843

26. Cyprinion macrostomum Heckel, 1843

27. Garra rufa (Heckel, 1843)

28. Garra variabilis (Heckel, 1843)

29. Hemigrammocapoeta elegans (Günther, 1868)

30. Iranocypris typhlops Bruun and Kaiser, 1944

31. Leuciscus cephalus (Linnaeus, 1758

32. Leuciscus lepidus (Heckel, 1843)

33. Pseudophoxinus persidis Coad, 1981

Family Cobitidae

1. Cobitis taenia Linnaeus, 1758

Family Balitoridae

1. Nemacheilus angorae Steindachner, 1897

2. Nemacheilus frenatus (Heckel, 1843)

3. Nemacheilus kermanshahensis Bănărescu and Nalbant, 1967

4. Nemacheilus malapterurus (Valenciennes in Cuvier and Valenciennes, 1846)

5. Nemacheilus smithi Greenwood, 1976

6. Nemacheilus tigris (Heckel, 1843)

Family Bagridae

1. Mystus pelusius (Solander in Russell, 1794)

Family Siluridae

1. Silurus triostegus Heckel, 1843

Family Sisoridae

1. Glyptothorax kurdistanicus (Berg, 1931)

2. Glyptothorax silviae Coad, 1981

Family Cyprinodontidae

1. Lebias ?? Aphanius dispar (Rüppell, 1829)

2. Aphanius mento (Heckel, 1843)

3. Aphanius punctatus (Heckel, 1849)

4. Aphanius vladykovi Coad, 1988

Family Mugilidae

1. Liza abu (Heckel, 1843)

2. Mugil cephalus Linnaeus, 1758

Family Gobiidae

1. Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837

2. Glossogobius giuris (Hamilton-Buchanan, 1822)

3. Periophthalmus waltoni Kouman, 1955

Family Mastacembelidae

1. Mastacembelus mastacembelus (Banks and Solander in Russell, 1794)

Namak Lake

Family Cyprinidae

1. Alburnoides bipunctatus (Bloch, 1782)

2. Barbus mursa (Gueldenstaedt, 1773)

3. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

4. Capoeta buhsei Kessler, 1877

5. Capoeta capoeta (Gueldenstaedt, 1773)

6. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

7. Leuciscus cephalus (Linnaeus, 1758

8. Leuciscus lepidus (Heckel, 1843)

Family Balitoridae

1. Nemacheilus bergianus Derzhavin, 1934

2. Nemacheilus malapterurus (Valenciennes in Cuvier and Valenciennes, 1846)

Family Salmonidae

1. Salmo trutta Linnaeus, 1758

Family Cyprinodontidae

1. Lebias punctata ?? Aphanius punctatus (Heckel, 1849)

Kavir

Family Cyprinidae

1. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

2. Capoeta capoeta (Gueldenstaedt, 1773)

3. Schizothorax pelzami Kessler, 1870

Bejestan

Family Cyprinidae

1. Capoeta capoeta (Gueldenstaedt, 1773)

2. Capoeta fusca Nikol'skii, 1897

3. Garra rossica (Nikol'skii, 1900)

Family Balitoridae

1. Nemacheilus kessleri Günther, 1889

Esfahan

Family Cyprinidae

1. Alburnoides bipunctatus (Bloch, 1782)

2. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

3. Capoeta capoeta (Gueldenstaedt, 1773)

4. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

5. Chalcalburnus mossulensis (Heckel, 1843)

6. Leuciscus lepidus (Heckel, 1843)

Family Cyprinodontidae

1. Lebias punctata ??Aphanius punctatus (Heckel, 1849)

Kerman-Na'in

Family Cyprinidae

1. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

2. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

Family Balitoridae

1. Nemacheilus sargadensis Nikol'skii, 1899

Lut

Family Cyprinidae

1. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

2. Capoeta fusca Nikol'skii, 1897

3. Cyprinion watsoni (Day, 1872)

4. Garra rossica (Nikol'skii, 1900)

Family Balitoridae

1. Nemacheilus bampurensis Nikol'skii, 1899

2. Nemacheilus sargadensis Nikol'skii, 1899

Sistan

Family Cyprinidae

1. Capoeta fusca Nikol'skii, 1897

2. Crossocheilus adiscus (Annandale, 1919)

3. Cyprinion watsoni (Day, 1872)

4. Garra persica Berg, 1913

5. Garra rossica (Nikol'skii, 1900)

6. Schizocypris altidorsalis Bianco and Banarescu, 1982

7. Schizopygopsis stoliczkai Steindachner, 1866

8. Schizothorax intermedius (McClelland, 1842)

9. Schizothorax zarudnyi (Nikol'skii, 1897)

Family Balitoridae

1. Nemacheilus kessleri Günther, 1889

2. Nemacheilus rhadinaeus Regan, 1906

3. Nemacheilus stoliczkai (Steindachner, 1866)

4. Nemacheilus tenuis Day, 1876

Kor River

Family Cyprinidae

1. Alburnoides bipunctatus (Bloch, 1782)

2. Barbus barbulus Heckel, 1849

3. Barbus luteus (Heckel, 1843)

4. Barbus pectoralis Heckel, 1843

5. Capoeta aculeata (Valenciennes in Cuvier and Valenciennes, 1844)

6. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

7. Chalcalburnus mossulensis (Heckel, 1843)

8. Chondrostoma regium (Heckel, 1843)

9. Cyprinion tenuiradius Heckel, 1849

10. Garra rufa (Heckel, 1843)

11. Pseudophoxinus persidis Coad, 1981

Family Cobitidae

1. Cobitis linea (Heckel, 1849)

Family Balitoridae

1. Nemacheilus malapterurus (Valenciennes in Cuvier and Valenciennes, 1846)

2. Nemacheilus persa (Heckel, 1849)

Family Cyprinodontidae

1. Lebias ?? Aphanius sophiae (Heckel, 1849)

Family Mastacembelidae

1. Mastacembelus mastacembelus (Banks and Solander in Russell, 1794)

Sirjan

Family Cyprinidae

1. Capoeta capoeta (Gueldenstaedt, 1773)

2. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

Family Balitoridae

1. Nemacheilus sargadensis Nikol'skii, 1899

Lake Maharlu

Family Cyprinidae

1. Barbus luteus (Heckel, 1843)

2. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

3. Chalcalburnus mossulensis (Heckel, 1843)

4. Cyprinion tenuiradius Heckel, 1849

5. Garra rufa (Heckel, 1843)

Family Balitoridae

1. Nemacheilus persa (Heckel, 1849)

Family Cyprinodontidae

1. Lebias punctata ?? Aphanius punctatus (Heckel, 1849)

?Liza abu

Gulf

Family Clupeidae

1. Tenualosa ilisha (Hamilton-Buchanan, 1822)

Family Cyprinidae

1. Barbus barbulus Heckel, 1849

2. Barbus grypus Heckel, 1843

3. Barbus kersin Heckel, 1843

4. Barbus luteus (Heckel, 1843)

5. Barbus pectoralis Heckel, 1843

6. Barilius mesopotamicus Berg, 1932

7. Capoeta barroisi (Lortet in Barrois, 1894)

8. Capoeta capoeta (Gueldenstaedt, 1773)

9. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

10. Chalcalburnus mossulensis (Heckel, 1843)

11. Chondrostoma regium (Heckel, 1843)

12. Cyprinion macrostomum Heckel, 1843

13. Cyprinion tenuiradius Heckel, 1849

14. Garra rufa (Heckel, 1843)

15. Pseudophoxinus persidis Coad, 1981

Family Cobitidae

1. Cobitis linea (Heckel, 1849)

Family Sisoridae

1. Glyptothorax silviae Coad, 1981

Family Cyprinodontidae

1. Lebias ?? Aphanius dispar (Rüppell, 1829)

2. Aphanius punctatus (Heckel, 1849)

Family Mugilidae

1. Liza abu (Heckel, 1843)

2. Mugil cephalus Linnaeus, 1758

Family Gobiidae

1. Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837

2. Glossogobius giuris (Hamilton-Buchanan, 1822)

3. Periophthalmus waltoni Koumans, 1955

Family Mastacembelidae

1. Mastacembelus mastacembelus (Banks and Solander in Russell, 1794)

Jaz Murian

Family Cyprinidae

1. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

2. Cyprinion watsoni (Day, 1872)

3. Garra persica Berg, 1913

4. Garra rossica (Nikol'skii, 1900)

Family Balitoridae

1. Nemacheilus bampurensis Nikol'skii, 1899

2. Nemacheilus sargadensis Nikol'skii, 1899

Family Cyprinodontidae

1. Lebias ?? Aphanius dispar (Rüppell, 1829)

Family Channidae

1. Channa gachua (Hamilton-Buchanan, 1822)

Mashkid

Family Cyprinidae

1. Aspidoparia morar (Hamilton-Buchanan, 1822)

2. Crossocheilus latius (Hamilton-Buchanan, 1822)

3. Cyprinion watsoni (Day, 1872)

4. Garra rossica (Nikol'skii, 1900)

Family Balitoridae

1. Nemacheilus kessleri Günther, 1899

2. Nemacheilus sargadensis Nikol'skii, 1899

Family Cyprinodontidae

1. Lebias ?? Aphanius dispar (Rüppell, 1829)

Hormuz

Family Clupeidae

1. Tenualosa ilisha (Hamilton-Buchanan, 1822)

Family Chanidae

1. Chanos chanos (Forsskål, 1775)

Family Cyprinidae

1. Barbus grypus Heckel, 1843

2. Barbus luteus (Heckel, 1843)

3. Capoeta buhsei Kessler, 1877

4. Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842)

5. Chalcalburnus mossulensis (Heckel, 1843)

6. Cyprinion watsoni (Day, 1872)

7. Garra persica Berg, 1913

8. Pseudophoxinus persidis Coad, 1981

Family Cobitidae

1. Cobitis linea (Heckel, 1849)

Family Cyprinodontidae

1. Lebias ?? Aphanius dispar (Rüppell, 1829)

2. Aphanius ginaonis (Holly, 1929)

Family Cichlidae

1. Iranocichla hormuzensis Coad, 1982

Family Mugilidae

1. Mugil cephalus Linnaeus, 1758

Family Gobiidae

1. Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837

2. Glossogobius giuris (Hamilton-Buchanan, 1822)

3. Periophthalmus waltoni Kouman, 1955

Makran

Family Cyprinida

1. Aspidoparia morar (Hamilton-Buchanan, 1822)

2. Crossocheilus latius (Hamilton-Buchanan, 1822)

3. Cyprinion watsoni (Day, 1872)

4. Garra rossica (Nikol'skii, 1900)

Family Balitoridae

1. Nemacheilus bampurensis Nikol'skii, 1899

Family Cyprinodontidae

1. Lebias ? Aphanius dispar (Rüppell, 1829)

Family Mugilidae

1. Mugil cephalus Linnaeus, 1758

Family Gobiidae

1. Boleophthalmus dussumieri Valenciennes in Cuvier and Valenciennes, 1837

2. Glossogobius giuris (Hamilton-Buchanan, 1822)

3. Periophthalmus waltoni Koumans, 1955

Summary of Native Distributions (% in parentheses)

Drainage Basin Families Genera Species
All Iran 25(100) 68(100) 155(100)
Lake Orumiyeh 4(16) 10(15) 12(8)
Caspian Sea 16(64) 43(63) 70(45)
Hari River 4(16) 10(15) 12(8)
Tigris River 12(48) 28(41) 56(36)
Namak Lake 4(16) 7(10) 12(8)
Kavir 1(4) 2(3) 3(2)
Bejestan 2(8) 3(4) 4(3)
Esfahan 2(8) 5(7) 7(5)
Kerman Na'in 2(8) 2(3) 3(2)
Lut 2(8) 4(6) 6(4)
Sistan 2(8) 8(12) 13(8)
Kor River 5(20) 12(18) 16(10)
Sirjan 2(8) 2(3) 3(2)
Lake Maharlu 3(12) 7(10) 7(5)
Gulf 8(32) 18(26) 26(17)
Jaz Murian 4(16) 6(9) 8(5)
Mashkid 3(12) 6(9) 7(5)
Hormuz 8(32) 15(22) 18(12)
Makran 5(20) 10(15) 10(6)


Threatened Species

Under construction.


Types

The nominal species originally described from Iran are summarised below by family, by year of description and then alphabetically. Some taxa have no type material (e.g. Eugen Keyserling did not bring specimens back to Europe) or type material has not been located despite a thorough search and is marked with an asterisk (*). Types whose existence is uncertain are indicated by a question mark (?). Type material examined by me is indicated by a plus sign (+) and type material not seen by me by an octothorpe (#). The current allocation of taxa is given in parentheses, those without comment being valid; material of uncertain status is marked with a tilde (~).

Details on these fishes can be found in the Species Accounts, where other type material examined for its importance in studying the Iranian ichthyofauna is listed. See also Eschmeyer's Catalog of Fishes (www.calacademy.org/research/ichthyology/catalog/).

Family Clupeidae

# Caspialosa caspia var. persica Il'in, 1927 (= Alosa caspia (Eichwald, 1838))

# Caspialosa knipowitschi Il'in, 1927 (= Alosa caspia (Eichwald, 1838))

# Clupea caspio-pontica var. grimmi Borodin, 1904 (= Alosa braschnikowii (Borodin, 1904))

Family Cyprinidae

*# Cyprinus persa Gmelin, 1774 (nomen nudum = Vimba vimba (Linnaeus, 1758))

*# Cyprinus bulatmai Hablizl, 1783 (= Barbus capito (Güldenstaedt, 1773))

?# Cyprinus chalybatus Pallas, 1814 (= Barbus capito (Güldenstaedt, 1773))

+ Alburnus capito Heckel, 1843 (= Alburnus mossulensis Heckel, 1843)

+ Chondrostoma aculeata Valenciennes in Cuvier and Valenciennes, 1844 (Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844))

# Leuciscus albuloides Valenciennes in Cuvier and Valenciennes, 1844 (= Alburnus chalcoides (Güldenstaedt, 1772))

+ Leuciscus maxillaris Valenciennes in Cuvier and Valenciennes, 1844 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus caudimacula Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus iblis Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus megacephalus Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Alburnus schejtan Heckel, 1849 (= Alburnus mossulensis Heckel, 1843)

+ Barbus barbulus Heckel, 1849

+ Cyprinion tenuiradius Heckel, 1849

+ Discognathus crenulatus Heckel, 1849 (= Garra rufa (Heckel, 1843))

+ Scaphiodon amir Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Scaphiodon macrolepis Heckel, 1849 (= Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844))

+ Scaphiodon niger Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Scaphiodon saadii Heckel, 1849 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

+ Systomus albus var. alpina Heckel, 1849 (= Barbus luteus (Heckel, 1843))

*#~ Alburnus maculatus Keyserling, 1861

*# Scaphiodon chebisiensis Keyserling, 1861 (Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

*# Scaphiodon gracilis Keyserling, 1861 (= Capoeta capoeta (Güldenstaedt, 1773))

*# Scaphiodon rostratus Keyserling, 1861 (= Capoeta damascina (Valenciennes in Cuvier and Valenciennes, 1842))

?# ~ Barbus miliaris De Filippi, 1863

+~ Alburnus doriae De Filippi, 1865

+ Schizothorax pelzami Kessler, 1870

+ Capoeta buhsei Kessler, 1877

+ Aspiostoma zarudnyi Nikol'skii, 1897 (= Schizothorax zarudnyi (Nikol'skii, 1897))

+ Capoeta fusca Nikol'skii, 1897

+ Capoeta gibbosa Nikol'skii, 1897 (= Capoeta capoeta (Güldenstaedt, 1773))

+ Capoeta nudiventris Nikol'skii, 1897 (= Capoeta fusca Nikol'skii, 1897)

+ Abramis urmianus Günther, 1899 (= Acanthalburnus urmianus (Günther, 1899))

+ Barbus bampurensis Nikol'skii, 1899 (= Cyprinion watsoni (Day, 1872))

+ Cyprinion kirmanense Nikol'skii, 1899 (= Cyprinion watsoni (Day, 1872))

+ Gobio persa Günther, 1899 (= Romanogobio persus (Günther, 1899))

+ Leuciscus gaderanus Günther, 1899 (= Petroleuciscus ulanus (Günther, 1899))

+ Leuciscus ulanus Günther, 1899 (= Petroleuciscus ulanus (Günther, 1899))

+ Cirrhina afghana var. nikolskii Berg, 1905 (= Cyprinion watsoni (Day, 1872))

+ Discognathus rossicus var. nudiventris Berg, 1905 (= Garra rossica (Nikol'skii, 1900))

+ Scaphiodon macmahoni Regan, 1906 (= Cyprinion watsoni (Day, 1872))

+ Garra persica Berg, 1913

+ Discognathus adiscus Annandale, 1919 (= Crossocheilus latius (Hamilton-Buchanan, 1822))

+ Discognathus phryne Annandale, 1919 (= Garra rossica (Nikol'skii, 1900))

+ Alburnus atropatenae Berg, 1925 (= Alburnus atropatenae Berg, 1925)

?# Alburnus alburnus hohenackeri natio persicus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Alburnus pseudospirlinus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Alburnus striatus Petrov, 1926 (= Alburnus alburnus (Linnaeus, 1758))

?# Rutilus rutilus caspicus natio knipowitschi Pravdin, 1927

+ Barbus baschakirdi Holly, 1929 (= Cyprinion watsoni (Day, 1872))

*# Barbus kessleri Derzhavin, 1929 (Barbus miliaris De Filippi, 1863)

*# Varicorhinus bergi Derzhavin, 1929 (= Capoeta aculeata Valenciennes in Cuvier and Valenciennes, 1844)

*# Varicorhinus nikolskii Derzhavin, 1929 (= Capoeta buhsei Kessler, 1877)

+ Barilius mesopotamicus Berg, 1932

?# Leuciscus cephalus orientalis natio ardebilicus Barach, 1934

+ Iranocypris typhlops Bruun and Kaiser, 1944

+ Chalcalburnus chalcoides iranicus Svetovidov, 1945 (= Alburnus chalcoides (Güldenstaedt, 1772))

+ Garra rufa gymnothorax Berg, 1949 (Garra rufa (Heckel, 1843))

+ Oreinus anjac Fowler and Steinitz, 1956 (Schizothorax zarudnyi (Nikol'skii, 1897))

+ Schizothorax schumacheri Fowler and Steinitz, 1956 (= Schizothorax intermedius McClelland, 1842)

+ Capoeta barroisi persica Karaman, 1969

+ Schizothorax pelzami iranicus Karaman, 1969 (= Schizothorax pelzami Kessler, 1870)

+ Pseudophoxinus persidis Coad, 1981 (Petroleuciscus persidis (Coad, 1982))

?# Barbus parieschanica Wossughi, Khoshzahmat and Etemadfar, 1982 (= Barbus luteus (Heckel, 1843))

+ Capoeta barroisi mandica Bianco and Banarescu, 1982

+ Capoeta capoeta intermedia Bianco and Banarescu, 1982

+ Chondrostoma cyri orientalis Bianco and Banarescu, 1982 (= Chondrostoma regium (Heckel, 1843))

+ Schizocypris altidorsalis Bianco and Banarescu, 1982

+ Barbus sublimus Coad and Najafpour, 1997

Family Cobitidae

+ Acanthopsis linea Heckel, 1849 (= Cobitis linea (Heckel, 1849))

# Cobitis aurata De Filippi, 1863 (= Sabanejewia aurata (De Filippi, 1863))

Family Balitoridae

+ Cobitis persa Heckel, 1849 (= Barbatula persa (Heckel, 1849))

+ Nemacheilus bampurensis Nikol'skii, 1899 (= Schistura bampurensis (Nikol'skii, 1899)

+ Nemacheilus sargadensis Nikol'skii, 1899 (= Schistura sargadensis (Nikol'skii, 1899)

+ Nemachilus rhadinaeus Regan, 1906 (= Paracobitis rhadinaea (Regan, 1906))

+ Nemachilus macmahoni Chaudhuri, 1909 (= Paracobitis rhadinaea (Regan, 1906))

+ Nemachilus bergianus Derzhavin, 1934 (= Barbatula bergiana (Derzhavin, 1934))

+ Noemacheilus kermanshahensis Bănărescu and Nalbant, 1967 (= Barbatula kermanshahensis (Bănărescu and Nalbant, 1967))

+ Nemacheilus smithi Greenwood, 1976 (= Paracobitis smithi Greenwood, 1976))

+ Orthrias farsicus Nalbant and Bianco, 1998 (= Barbatula farsica (Nalbant and Bianco, 1998))

+ Paracobitis iranica Nalbant and Bianco, 1998

+ Paracobitis vignai Nalbant and Bianco, 1998

+ Schistura nielseni Nalbant and Bianco, 1998

? Seminemacheilus tongiorgii Nalbant and Bianco, 1998

Family Sisoridae

+ Glyptosternum kurdistanicum Berg, 1931 (= Glyptothorax kurdistanicus (Berg, 1931))

+ Glyptothorax silviae Coad, 1981

Family Cyprinodontidae

+ Lebias crystallodon Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

+ Lebias punctatus Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

+ Lebias sophiae Heckel, 1849 (= Aphanius sophiae (Heckel, 1849))

#~ Cyprinodon blanfordii Jenkins, 1910

# Cyprinodon persicus Jenkins, 1910 (= Aphanius persicus (Jenkins, 1910))

#~ Cyprinodon pluristriatus Jenkins, 1910

+ Cyprinodon ginaonis Holly, 1929 (= Aphanius ginaonis (Holly, 1929)

+ Aphanius vladykovi Coad, 1988

+ Aphanius isfahanensis Hrbek, Keivany and Coad, 2007

Family Cichlidae

+ Iranocichla hormuzensis Coad, 1982

Family Mugilidae

+ Liza abu zarudnyi Berg, 1949

Family Gobiidae

# Gobius semipellucidus Kessler, 1877 (= Proterorhinus marmoratus (Pallas. 1814))


F. Glossaries

Geographical Glossary

The following glossary of geographical and other terms is mostly in Farsi (which includes words taken from Turkic, Kurdish and Arabic) with a few Russian words, all of which appear on maps, in the literature and in the text of the "Freshwater Fishes of Iran" web site. They may be of help to those unfamiliar with these languages and avoid tautologies such as Safidrud River.

There are various diacritical marking systems for Farsi but these do not always transfer accurately across platforms, appearing as strange symbols or gaps in text. I have eschewed the use of these as being irrelevant for native speakers of Farsi and too confusing for those unfamiliar with this language.

ab = water, intermittent stream, stream, spring, lake, well

ab-bandan = shallow, freshwater reservoir on the Caspian plain used for duck hunting in winter and water storage for irrigation in the dry summer

abad = a suffix indicating an inhabited place

ab-e garm = hot spring

ab ambar = cistern

ab anbar = cistern

abshur rud = salt river, common name of salty rivers

anbar = tank

ateshkade = fire-temple (archaeological feature)

av = stream

`ayn = spring

bagh = garden

bahr = sea

Bahr-e Khazar = Caspian Sea

baksh= municipality

band = dam, reservoir, lake, mountain range (old dams for water storage - see sadd for modern dams)

bandar = port, harbour, anchorage, bay

bankari = constructing temporary weirs for water diversion

bar andaz = halting place

barm = marsh, lake, pond

batlaq = marsh, swamp

berkeh = tank, pool, cistern

biaban = desert (also the name of the coastal plain south of the Minab River to the cape of Ra's al-Kuh in Hormozgan Province)

birkat = pool, well, marsh, lake

borj = fort, tower

botlaq = marsh, swamp

caviar = sturgeon eggs as food

çay = stream

centner = 100 kg (used in Russian texts as a measure of commercial catches; sometimes given as 50 kg elsewhere but internal evidence in Russian papers indicates 100 kg is correct)

chah = well, spring, cistern

chai = stream

cham = stream, gorge

chashmeh = spring, well

chay = stream

cheng = hill, mountain, promontory

cheshmeh = spring, well

dag = mountain

dagh = mountain

dahaneh = section of a stream, gorge, pass, defile, water gap

damagheh = cape, promontory

damgah = an artificial, freshwater wetland, maintained primarily as a duck-hunting area but also used for irrigation during the dry summer months

daqq = salt flat, salt depression, salt waste, marsh, intermittent salt lake

darband = gorge or pass

darreh = stream, valley, gorge, ravine

darya = sea, stream, intermittent stream, channel

daryacheh = lake, marshy lake, stream

Darya-ye Mazandaran = Caspian Sea

dasht = plain, desert, steppe, depression, upland, open country, field; usually dry desert with a firm base of pebbles or silts

deh = village

dehkadeh = village

dez = fortress

echbel = eggs of fishes other than sturgeons as food

emamzadeh = tomb, shrine

eskeleh = jetty

estakhr = pool

gadik, gaduk = pass

galal = stream

gardan, gardaneh = pass

godar = pass

garmsir = hot country, winter quarters in the lowlands

ghadamgah = a religious site; usually no fishing allowed.

gharb(i) = west(ern), as in the province Azarbayjan-e Gharbi)

godar = pass

göl = lake, marsh, swamp

gölü = lake, marsh, swamp

gowd = depression

hammam = bath

hamun = marshy lake, salt waste

hawr = marsh, lake

hesar = fort, castle

hor = marsh

howr = marsh

howz = tank, cistern, pond, pool, lake, reservoir, spring

il'men = a shallow, flood-plain lake heavily overgrown with reeds and rushes (Russian)

ishan = hill

istgah = railway station

jabal = hill, mountain

jangal = forest

jar = stream

jazirat = island

jazireh = island

jebal = hill, mountain

jehil = lake

jolgeh = plain

jonub(i) = south(ern)

ju = stream, irrigation channel

jub, jube = irrigation channel, watercourse, gutter, ditch

juy = stream, watercourse

kal = stream, intermittent stream

kalleh = peak

kamar = hill, mountain, ridge

kani = well

karavansara = caravansary

karez = underground irrigation channel

kaur = stream

kavir = salt waste, salt desert, marsh; usually a salt crust over silt deposits which can be fatal mires of slimy mud (= playa)

khalij = bay, gulf

Khalij-e Fars = Persian Gulf

kharabeh = ruins

khirr = stream

khowr = inlet, stream, channel, bay, bight, tidal creek, estuary

khwar = stream

kizil ala = brown trout (see also qezelala)

kotal = mountain pass

kowr = stream

kuh = mountain, range, hill, peak, ridge, spur

kuhha = mountains, range, hills

liman = a brackish bay of the sea, usually at a river mouth, sometimes cut off from the sea but still brackish; also an estuary (Russian, mordab in Farsi)

lut = desert

mahur = hill

mandah = stream

markazi = central, as in Markazi Province

masjed = mosque

mordab = lagoon, backwater, creek, swamp, stagnant water (literally dead water, now replaced by talab)

nahr = river, stream, canal, docking basin

naizar = reed swamp (Sistan)

namak = salt; usually a salt lake with open water or a salt crust but without much mud

namaksar = salt waste

naveh = stream

nawah = stream

nehri = stream

neizar = reed swamp in Sistan

ostan = province, governorate-general

ozero = lake

pal = hill, mountain

paskuh = mountain range

pereval = pass

poshteh = hill, mountain

qabr = tomb

qabrestan = cemetery

qal'at = fort

qal'eh = fortress

qanat = underground irrigation channel; an adit shaft

qasr = fort

qolleh = hill, mountain, peak

ramlat = sandy area

ra's = cape, point, promontory

reka = river

reshteh = mountain, mountain range, hill, spur

reshteh kuh = mountain range

rig = sand area, dunes

riz ab = stream

roga = outflow (in the Enzeli or Anzali Mordab)

rud = river, stream, intermittent stream

rudbar = valley drained by a river with flowing water; place watered by many streams

rudkhaneh = river, river bed, watercourse, intermittent stream

rusta = village, inhabited place

sabkhat = salt marsh, lake

sadd = dam, reservoir (used for modern dams)

saddi qanat = a qanat drawing water from a dam

sahel = coast, beach, shore

sar = cape

sarab = spring (in western Iran), literally "beginning water"

saray = caravansaray

sardsir = cold country, summer quarters in the highlands

sarhadd = frontier

sazhen = a marine sazhen equals 1.83 m (Russian)

selseleh = mountain range, mountains

shahr = town, city

shahrdari = municipality

shahrestan = district

shahzadeh = shrine

shamal(i) = north(ern)

sharq(i) = east(ern), as in the province Azarbayjan-e Sharqi

shatt = large river, bank of a river, stream

saydgah = fishing station, as along the Caspian coast

shebh-e jazireh = peninsula

shekasteh = hill, mountain

shil = a wooden barrier erected across a river for catching fish; hence shilat (in Gilaki, the Persian dialect of Gilan)

shilat = fisheries company; Sherkat Shilat = Northern Fisheries Company concerned with the Caspian Sea; Shilat Jonub = Southern Fisheries Company concerned with the Persian Gulf

shur = salt, brackish, stream

shurab = salt water

shurehzar = salt stream, salt marsh

su = water, stream

suyu = stream

talab = more modern version of mordab.

tall = hill, mountain, spur

tang = pass

tangeh = valley

tappeh = hill, mountain, mound

tell = hill

tepe = hill, mound

vareh = a small dam

vilayet = province

ziarat = shrine


Ichthyological Glossary

Technical terms used in ichthyology and in this work are in the Dictionary of Ichthyology and can also be accessed through the Main Page of www.briancoad.com.


G. Bibliography

This section contains all the papers and books referred to in the text of the web site on Freshwater Fishes of Iran but also attempts to be a bibliography on the freshwater fishes of Iran and their environment. A partial version of this bibliography was first posted on 10 September 1996 and maintained by Yazdan Keivany, then at the University of Alberta, Edmonton.

The large number of articles listed here, the length of many, their limited availability, and costs involved make it impossible to loan xeroxes except in restricted cases. Please use your interlibrary loan service.

There has been a rapid increase in studies on fishes of Iran, starting in the 1990s. Prior to 1900, this bibliography lists less than 100 publications relevant to this work, many not strictly on Iranian fishes. On a decadal basis, it is only in the 1960s that publications exceed 100 and by the 1990s are an order of magnitude larger. Many articles are available as an abstract at www.sid.ir/En/Index.asp.

Use of diacritical marks in various languages in this Bibliography may not always transfer across platforms. As a consequence, some diacritical marks can appear as gibberish letters (with apologies to colleagues in eastern Europe and Turkey).

Note that articles published in the "Journal of Ichthyology" are translations from the original Russian journal "Voprosy Ikhtiologii" although pagination (and sometimes issue number) are not the same. Only "Journal of Ichthyology" pagination is cited here but readers may wish to avail themselves of the Russian versions of these articles (however some earlier issues of "Voprosy Ikhtiologii" are only available in Russian). Prior to 1970 the English translation was called "Problems of Ichthyology". Year of publication as marked on the cover of each issue was the one cited here although internal evidence indicates translated issues were often published in the following year, a delay presumably because of translation time required. The date thus used is the Russian date of publication which is the relevant one for the few taxon descriptions appearing in this journal.

There is a moderately extensive grey literature on Iran, mostly manuscript reports of the Department of the Environment, Tehran. Some of these are in English and were written by Peace Corps workers in the 1960s and 1970s. I have cited all those I came across but the listings are by no means exhaustive. Additionally, magazines and newspapers in Iran carry articles on fishes. Regretfully, a complete survey of these was not possible.

The internet now carries English translations of Iranian newspapers and press releases. The URL (Universal Resource Locator) for these and other sites having information on Iran are cited directly in the text and not in this bibliography. Note that links often change or are no longer functional and these textual references serve only to document a source. Generally the date when they were downloaded is given. They may still be accessible but are not hot linked in the text. Some papers cited in this Bibliography have only been seen in their online version and pagination of the original, printed version is unknown.

Transliteration of personal names into English from other languages, notably Farsi, often gives variant spellings. When I am aware that the same person is meant, the original journal spelling is given followed by (sic) to indicate that the author may appear under another spelling. The same (sic) is also used to denote misspellings in titles. Note that family names in Farsi may appear in two forms, a single name or a double-barreled name (separated or hyphenated), and these are variously used in papers. Personal names may be abbreviated to a single letter when appearing in English but sometimes appears as two letters to accord with the Persian alphabet (e.g. there are single letters s and sh).

The "Iranian Journal of Fisheries Sciences" (from 7(2) 1998 onwards) also appeared earlier as the "Iranian Fisheries Scientific Journal" (3(2) 1994 to 7(1) 1998), and earlier still as "Iranian Fisheries Journal", "Iranian Fisheries Scientific Bulletin" and "Iranian Fisheries Bulletin", is published in Farsi with English abstracts. It sometimes appears as the "Iranian Scientific Fisheries Journal" in Iranian abstracting journals and websites. The "Iranian Journal of Fisheries Sciences" (starting in January 1999 with volume 1(1)) is published in English with Farsi abstracts. The Iranian Fisheries Research Organization (formerly Iranian Fisheries Research and Training Organization) publishes these journals and has a website with lists of publications.

The website http://www.sid.ir/En/Index.asp has lists of publications in Iranian journals and abstracts, both in English, as well as in Farsi.

Students of Iranian fishes should also consult the Freshwater Fishes of Iraq website for its Bibliography as this includes additional and complementary studies on species shared with Iran.

A

Abaee, S. 2001. Rapid assessment of point sources pollution in Iranian part of Caspian Sea area using GIWA methodology. Caspian Environment Programme, Baku, Azerbaijan and Tehran. 8 pp., 3 figs., appendices A-B.

Abakumov, V. A. 1960. The damage done by lampreys to fish stocks. Fisheries Research Board of Canada Translation Series, 274:1-2.

Abassali Zadeh, A. 2003. Effects of ablading of sensory barbels on growth of Huso huso (young stage). Iranian Journal of Fisheries Sciences, 12(3):193-200. In Farsi.

Abaychi, J. and Al-Saad, H. T. 1988. Trace elements in fish from the Arabian Gulf and the Shatt al-Arab River, Iraq. Bulletin of Environmental Contamination and Toxicology, 40:226-232.

Abaychi, J. K., Darmoian, S. A. and DouAbul, A. A. Z. 1988. The Shatt al-Arab River: A nutrient salt and organic matter source to the Arabian Gulf. Hydrobiologia, 166:217-224

Abbasi, H. 1974. Neur Lake experimental commercial fishery, 27 Mehr - 16 Aban. L-6-53, Job Progress Report, Department of the Environment, Tehran. 9 pp. MS.

Abbasi, H. 1975. Neur Lake commercial fishery, 29 Mehr - 19 Aban 1354. Job Progress Report, Fisheries and Aquatic Ecology Section, Department of the Environment, Tehran. MS (Abstract).

Abbasi, K. 2003. First record of black carp, Mylopharyngodon piceus from southern Caspian Sea (Iran). Iranian Journal of Fisheries Sciences, 12(2):139-146. In Farsi.

Abbasi, K. 2005. Studying alien fishes and macrocrustaceans distribution and their effects on rivers and wetlands of the Iranian basin of Caspian Sea. Borok II, International Workshop, Invasions of Alien Species in Holarctic, 27 September-1 October 2005, Borok (abstract).

Abbasi, K. 2006a. Identification and distribution of fish fauna in Shafarud River, Guilan Province. Iranian Journal of Fisheries Sciences, 15(2):73-86. In Farsi.

Abbasi, K. 2006b. Identification and distribution of fish fauna in Hevigh River (Guilan Province). Iranian Journal of Biology, 18(4):370-382. In Farsi.

Abbasi, K., Keyvan, A. and Ahmadi, M. R. 2004. Morphometric and meristic characteristics of Vimba vimba persa in Sefidrud River. Iranian Journal of Fisheries Sciences, 13(1):61-76. In Farsi.

Abbasi, K., Keyvan, A. and Ahmadi, M. R. 2005. Studying natural reproduction, spawning grounds and spawning period of Vimba vimba persa population in Sefid-Roud River, Guilan Province in north Iran. Iranian Journal of Fisheries Sciences, 14(3):113-126. In Farsi.

Abbasi, K., Moraadkha, S. and Sarpanah, A. 2007. Identification and distribution of fish fauna in Siahdarvishan river (Anzali Wetland basin). Pajouhesh va Sazandegi, 74:27-39. In Farsi.

Abbasi, K. and Sabkara, J. 2004a. Studying food items of Leuciscus ulanus, an endemic fish species in Iran. The 1st Congress on Animal and Aquatic Sciences, University of Tehran, 31 August - 2 September 2004, p. 492-495.

Abbasi, K. and Sabkara, J. 2004b. Studying some biological characteristics of Leuciscus ulanus (Cyprindae), an endemic fish and little-known in Iran. 2nd International Conference on Applied Biology, Mashad, 29-30 September 2004, p. 29.

Abbasi, K. and Sabkara, J. 2004c. Studying Alosa caspia caspia (Caspian shad) diet in southeastern Caspian Sea (Mazandaran and Golestan provinces). Iranian Journal of Biology, 17(3):272-290. In Farsi.

Abbasi, K., Salavatian, S. M. and Abdollapoor, H. 2005. Investigating fish diversity and distribution in the Mahabad-Chai River of the Lake Urmia basin, north-western Iran. Iranian Journal of Fisheries Sciences, 13(4):75-94. In Farsi.

Abbasi, K. and Sarpanah, A. 2001. Fish fauna investigation in Aras Reservoir and its Iranian tributaries. Iranian Journal of Fisheries Sciences, 10(2):41-62. In Farsi.

Abbasi, K., Valipour, A., Talebi Haghighi, D., Sarpanah, A. and Nezami, Sh. 1999. Atlas of Iranian Fishes. Gilan Inland Waters. Gilan Fisheries Research Centre, Rasht. vi + 113 pp. In Farsi.

Abbasi, K. and Valipour, A. R. 2006. Studying the Silurus glanis Linnaeus, 1758 food items in Anzali lagoon. Pajouhesh va Sazandegi, 66:14-24. In Farsi.

Abbasian, G-R. 1997. Fisheries of Iran: Development possibilities. INFOFISH International, Kuala Lumpur, 2:14-18.

Abbasov, G. S. 1980. The ichthyofauna of the main freshwater bodies of Azerbaidzhan. Journal of Ichthyology, 20(4):140-142.

Abbaspour, M. and Sabetraftar, A. 2005. Review of cycles of drought and indices of drought and their effect on water resources, ecological, biological, agricultural, social and economical issues in Iran. International Journal of Environmental Studies, 62(6):709-724.

Abdi, K. 1999a. Survey on the four new species of parasitic leeches of Mahabad's Lake Dam in Iran. In: 26th World Veterinary Congress, Lyon, France, 23-26 September 1999 (abstract).

Abdi, K. 1999b. Survey on the crustacean parasites of fishes of Mahabad's reservoir dam with new records of Tracheliastes polycolpus in Iran. In: 26th World Veterinary Congress, Lyon, France, 23-26 September 1999 (abstract).

Abdi, K., Mobedi, I. and Rostamzad, R. 2006. The first outbreak of acanthocephalosis in rainbow trout fish farms of Iran. Pajouhesh va Sazandegi, 71:9-12. In Farsi.

Abdolhay, H. 1996. Aquaculture status and development in the Islamic Republic of Iran. Paper presented to the Working Group on Aquaculture, Indian Ocean Fishery Commission Committee for the Development and Management of the Fishery Resources of the Gulf, Cairo, Egypt, 1-3 October, 1996.

Abdolhay, H. 1997a. Artificial reproduction of fish for stock enhancement in the Caspian Sea. Seventh Conference of Shilat, Responsible Fisheries, 17-18 February 1997, Tehran. p. 187-207. In Farsi.

Abdolhay, H. 1997b. Fingerling production and stock enhancement of sturgeon in south of Caspian Sea. Third International Symposium on Sturgeon, Piacenza, Italy, 8-11 July 1997, Book of Abstracts (poster).

Abdolhay, H. and Baradaran Tahori, H. 1998. Review of reproduction of sturgeon from 1991-1997 in Iranian hatchery. First National Symposium on Sturgeon, Rasht, 15-16 December 1998, p. 107-124. In Farsi.

Abdolhay, H. and Baradaran Tahori, H. 1999. Fingerling production and stock enhancement of sturgeon in south of Caspian Sea. Journal of Applied Ichthyology, 15(4-5):298.

Abdolhay, H., Baradaran Tahouri, H. and Amini, R. 2006. An audit of sturgeon reproduction in Iran over the years 1998-2002. Iranian Scientific Fisheries Journal, 14(4):97-112. In Farsi.

Abdolhay, H. A. and Baradaran Tahori, H. 1999. Fingerling production and release for stock enhancement of sturgeon in the southern Caspian Sea: an overview. Journal of Applied Ichthyology, 22(supplement 1):125-131.

Abdoli, A. 1992. A new fish species in Mazandaran ab bandans. Abzeeyan, Tehran, 2(8):54-55. In Farsi.

Abdoli, A. 1993a. Threespine stickleback in the Caspian Sea. Abzeeyan, Tehran, 3(10 & 11):45. In Farsi.

Abdoli, A. 1993b. Fishes of Golestan National Park's river. Abzeeyan, Tehran, 4(3):32-33, XIII. In Farsi.

Abdoli, A. 1993c. Three-spined stickleback in southern basins of the Caspian Sea. Abzeeyan, Tehran, 4(5):10-13, III. In Farsi.

Abdoli, A. 1994a. Tadpole goby Benthophilus stellatus leobergius in Gorgan Bay. Abzeeyan, Tehran, 4(12):58-59. In Farsi.

Abdoli, A. 1994b. Ecological study of fish populations in Sardabrud River and Chalus River, Mazandaran, Iran. M.Sc. Thesis, University of Tehran. iv + 94 pp, 28 figs, 20 tables.

Abdoli, A. 1998. An investigation of some biological characteristics and distribution of Pseudorasbora parva in freshwaters of Iran. Journal of Agricultural Sciences and Natural Resources, 8(1):33-40. In Farsi.

Abdoli, A. 2000. The Inland Water Fishes of Iran. Iranian Museum of Nature and Wildlife, Tehran. 378 pp. In Farsi.

Abdoli, A., Coad, B. and Naderi, M. 2000. First record of Rhinogobius similis, Gill 1859 in Iran. Iranian Journal of Fisheries Sciences, 9(1):73-76, 9. In Farsi.

Abdoli, A., Naderi, M., Aboo, M., Fazli, H. and Afraee, M. A. 1996. Spawning time and fecundity of the mullet, Liza auratus (sic) in south eastern part of the Caspian Sea. Abzeeyan, Tehran, 7(2):24-26, III. In Farsi.

Abdoli, A and Rahmani, H. 1999. The populations, food consumption and reproduction of Neogobius fluviatilis in a small stream of Madarso, Golestan National Park, North East of Iran. Seventh International Symposium on the Ecology of Fluvial Fishes, 10-13 May 1999, Lodz, Poland (poster).

Abdoli, A. and Rahmani, H. 2001. Food habits of two species of Gobiidae Neogobius fluviatilis, Neogobius melanostomus in the Madarsoo stream, Golestan National Park. Journal of Agricultural Sciences and Natural Resources, 8(1):3-15, 191. In Farsi.

Abdoli, A., Rahmani, H. and Rasooli, P. 2002. On the occurrence, diet and reproduction of Neogobius fluviatilis in Madarsoo stream, Golestan National Park, (north eastern Iran). Zoology in the Middle East, 26:123-128.

Abdoli, A., Rasooli, P. and Mostafavi, H. 2008. Length-weight relationships of Capoeta capoeta capoeta (Gueldenstaedt, 1772) in the Gorganrud River, south Caspian basin. Journal of Applied Ichthyology, 24(1):96-98.

Abdoli, A., Rasooli, P. and Soltaninasab, S. 2008. A contribution to the biology of Acanthalburnus urmianus (Günther, 1899) (Osteichthyes: Cyprinidae): an endemic fish of Iran. Zoology in the Middle East, 43:111-112.

Abdoli, A. and Skandari, S. K. 1999. Reproductive biology of khramulya (Capoeta capoeta gracilis) in Madarso stream in Golestan National Park. Journal of Agricultural Sciences and Natural Resources, 6(3):38-51. In Farsi.

Abdolmalaki, Sh. 2001. Catch statistic review of mullet fishery in Iranian coastal waters of the Caspian Sea. Iranian Journal of Fisheries Sciences, 9(4):39-52. In Farsi.

Abdolmalaki, Sh. 2004. Fishery status and stock assessment of silver carp (Hypophthalmichthys molitrix) in Mahabad Reservoir in fishing season of 1998-1999. Iranian Journal of Fisheries Sciences, 13(1):77-96. In Farsi.

Abdolmalaki, Sh. 2005. Stock assessment of the pick (sic) perch (Stizostedion lucioperca) in the southern coastal waters of the Caspian Sea. Iranian Journal of Fisheries Sciences, 13(4):95-110. In Farsi.

Abdolmalaki, Sh. 2006a. Stock assessment of bonyfishes in the Iranian coastal water of the Caspian Sea. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture, Bandar Anzali. http://en.ifro.ir (abstract).

Abdolmalaki, Sh. 2006b. Population dynamics of Caspian sea bream (Abramis brama orientalis) in Iranian coast of the Caspian Sea in 2000-2001. Pajouhesh va Sazandegi, 68:85-92. In Farsi.

Abdolmalaki, Sh. 2006c. Trends in stock fluctuations of Rutilus frisii kutum in the Caspian Sea. Iranian Journal of Fisheries Sciences, 15(2):87-100. In Farsi.

Abdolmalaki, Sh. and Ghaninezhad, D. 1999. A survey on releasing fingerlings and catching pikeperch (Stizostedion lucioperca) in Iranian coastal waters of the Caspian Sea. Proceedings of the First National Congress on Caspian Sea Bony Fish, 21-23 January 1999, Bandar Anzali, Gilan. In Farsi.

Abdolmalaki, Sh. and Psuty, I. 2007. The effects of stock enhancement of pikeperch (Sander lucioperca) in Iranian coastal waters of the Caspian Sea. ICES Journal of Marine Sciences, 64:8 pp. (advance copy).

Abdolmaleki, S. 1994. An investigation on the bentic (sic) macrofauna of the Anzali Lagoon. Iranian Fisheries Scientific Bulletin, (5):27-38, 4. In Farsi.

Abdurakhmanov, Yu. A. 1962. Ryby Presnykh vod Azerbaidzhana [Freshwater Fishes of Azerbaidzhan]. Akademii Nauk Azerbaidzhanskoi SSR, Institut Zoologii, Baku. 407 pp.

Abdurrahmanov, Y. A. 1966. Azarbaycanin Faunasi. VII Cild. Baliglar (Pisces). [The Fauna of Azerbaidzhan. VII. Fishes (Pisces)]. Zoologiya Institutu, Azarbaycan SSR Elmlar Akademiyasi, Baki. 224 pp. In Azerbaijani.

Abdurakhmanov, Yu. A. 1975. Transformation of the diadromous Kura shemaya Chalcalburnus chalcoides into a land-locked population in the Mingechaur Reservoir. Journal of Ichthyology, 15(2):189-196.

Abdurakhmanov, Yu. A., Kuliev, Z. M. and Agayarova, A. E. 1968. Materialy po Biologii i Raspredeleniyu Ryb u Azerbaidzhanskogo Poberezh'ya Srednego i Yuzhnogo Kaspiya [Materials on the biology and distribution of fishes at the Azerbaidzhan coast of the Central and Southern Caspian]. In: The Biology of the Central and Southern Caspian. Moscow. pp. 113-146.

Abdurakhmanov, Yu. I. and Kuliyev, Z. M. 1968. European eel in Azerbaidzhan waters. Problems of Ichthyology, 8:592-594. In English.

Abdusamadov, A. S. 1986. Biology of white amur, Ctenopharyngodon idella, silver carp, Hypophthalmichthys molitrix, and bighead, Aristichthys nobilis, acclimatized in the Terek region of the Caspian basin. Journal of Ichthyology, 26(4):41-49.

Abedian Kennari, A. M., Oveisipour, M. R. and Nazari, R. M. 2007. Effects of n3-HUFA enriched Daphnia magna on growth, survival, stress resistance, and fatty acid composition of larvae of Persian sturgeon (Acipenser persicus). Iranian Journal of Fisheries Sciences, 7(1):1-14.

Abell, R., Thieme, M. L., Revenga, C., Bryer, M., Kottelat, M., Bogutskaya, N., Coad, B., Mandrak, N., Contreras Balderas, S., Bussing, W., Stiassny, M. L. J., Skelton, P., Allen, G. R., Unmack, P., Naseka, A., Ng, R., Sindorf, N., Robertson, J., Armijo, E., Higgins, J. V., Heibel, T. J., Wikramanayake, E., Olson, D., López, H. L., Reis, R. E., Lundberg, J. G., Sabaj Pérez, M. H. and Petry, P. 2008. Freshwater Ecoregions of the World: A new map of biogeographic units for freshwater biodiversity conservation. BioScience, 58(5):403-413.

Abich, H. 1856. Vergleichende Chemische Untersuchungen der Wasser des Caspischen Meeres, Urmia- und Van-See's. Mémoires de l'Académie Impériale des sciences de Saint-Pétersbourg, sixième série, Sciences mathématiques et physiques, 7:57 pp., 2 plates.

Abou-Seedo, F., Dadzie, S., Al-Kanaan, K. and Sukumaran, J. V. 2003. Aspects of reproductive biology of the hermaphroditic yellowfin seabream, Acanthopagrus latus (Hottuyn (sic), 1782), in cages in Kuwait Bay. Zoology in the Middle East, 29:51-58.

Abtahi B., Esmaili Sari, A., Khodabandeh, S. and Seifabadi J. 2001. Ctenophores of the Caspian Sea and their socio-economic impacts. The Second International Iran and Russia Conference (Agriculture & Natural Resources), Moscow. Abstract, p. 164.

Abtahi, B., Faghiehzadeh, S., Omidbeigi, R., Shariefpour, A., Rassoli, A., Nazari, R. M. and Aghajanpour, M. 2002. The comparison of LC50 of clove essence and MS222 in Acipenser persicus, Oncorhynchus mykiss and Cyprinus carpio. Iranian Scientific Fisheries Journal, 11(3):1-12. In Farsi.

Abtahi, B. and Nazari, R. M. 2006. Comparison of antifungal therapeutic indices of formalin, malachite green and potassium permanganate on Persian sturgeon in farming conditions of north of Iran (Sari). Pajouhesh va Sazandegi, 67:42-49. In Farsi.

Abtahi, B., Soltani, M., Mohammadi Arani, M., Amin, G. and Khoshbavar-Rostami, H. 2003. Anaesthetic effects of clove essence (Eugenia caryophyllata) on fingerlings of Persian sturgeon (Acipenser persicus). Journal of Agricultural Sciences and Natural Resources, 10(1):161-172. In Farsi.

Abtahi, B. and Taghavi, H. 2006. Anatomical and histological study of ovary maturity stages of common kilka (Clupeonella delicatula). Pajouhesh va Sazandegi, 63:47-54. In Farsi.

Abtahi, B., Taghavi, H., Fazli, H. and Usefian, M. 2002. Fecundity and some biometrical characteristics of common kilka (Clupeonella delicatula) in Mazandaran Province. Iranian Journal of Marine Sciences, 1(2):1-9, 88. In Farsi.

Abuzyarov, Z. K. 1999. Problems of the Caspian Sea level predictions. TACIS (Technical Assistance to the Commonwealth of Independent States, European Union), Caspian Environment Programme, Baku, Azerbaijan. 17 pp.

Abzigostar. 1996. Iran fisheries sector study. Report by Abzigostar Consulting Engineering Company to Shilat, Tehran. 190 pp.

Abzigostar. 2004a. Report on fisheries activity in Zarivar Lake. Report by Abzigostar Consulting Engineering Company to Fisheries General Directorate, Kurdistan Province. 68 pp.

Abzigostar. 2004b. Report on environment in Zarivar Lake. Report by Abzigostar Consulting Engineering Company to Fisheries General Directorate, Kurdistan Province. 95 pp.

Abzigostar. 2004c. Report on limnology in Zarivar Lake. Report by Abzigostar Consulting Engineering Company to Fisheries General Directorate, Kurdistan Province. 100 pp.

Adamicka, P. 1984. On the construction of the Pontic cyprinid Pelecus cultratus (L.). Archiv für Hydrobiologie, 101(1/2):9-19.

Adams, C. G., Bayliss, D. D. and Whittaker, J. E. 1999. The terminal Tethyan event: A critical review of the conflicting age determinations for the disconnection of the Mediterranean from the Indian Ocean, p. 477-484. In: Whybrow, P. J. and Hill, A. (Eds.). Fossil Vertebrates of Arabia with Emphasis on the Late Miocene Faunas, Geology, and Palaeoenvironments of the Emirate of Abu Dhabi, United Arab Emirates. Yale University Press, New Haven and London. xxv + 523 pp., 40 pp.

Adams, R. M. 1962. Agriculture and urban life in early southwestern Iran. Science, 136(3511):109-122.

Adeli Mosabbab, Y. and Piri, Kh. 2005. Monitoring of release rnies (sic) of juvenile vobla (Rutilus rutilus) at the Iranian coast of the Caspian Sea with the purpose of restoring of its stocks, p. 15. In: Proceedings XIII International Conference on Fisheries Oceanology, Svetlogorsk, Kanliningrad region, September 12-17, 2005.

Adibmoradi, M and Sheibani, M. T. 2002. The histological study of mesencephal and cerebellum in Rutilus frisii kutum. Journal of the Faculty of Veterinary Medicine, Tehran, 57(2):15-18.

Adler, K. 1989. Contributions to the History of Herpetology. Society for the Study of Amphibians and Reptiles, Contributions to Herpetology, Oxford, Ohio, 5:1-202.

Admiralty Naval Staff. 1918. Geology of Mesopotamia and its borderlands. Intelligence Department, Admiralty Naval Staff, HMSO, London (I. D. 1177). 116 pp.

Afifi, H. 1966. A summary of method analysis used in the Zayandeh Rud river discharge study, pp. 213-226. In: Symposium on Hydrology and Water Resources Development held in Ankara, Turkey February 7 to 12, 1966, Central Treaty Organization, Ankara. 484 pp.

Afkhami, A. A. 1998. Disease and water supply: the case of cholera in 19th century Iran. Yale School of Forestry & Environment Studies Bulletin, 103:206-220.

Afkhami, M., Shariat, M., Jaafarzadeh, N., Ghadiri, H. and Nabizadeh, R. 2007. Regional water quality management for the Karun-Dez River basin, Iran. Water and Environment Journal, 21(3):192-199.

Afraei, M. A. 2006. Identifcation and appointment distribution Clupeonella (genus Alosa) in the South Caspian Sea coastal. Iranian Fisheries Research Organization, Agriculture Research and Education Organization, Ministry of Jihad-e-Agriculture, Bandar Anzali. http://en.ifro.ir (abstract).

Afraei, M. A., Fazli, H. and Moslemi, M. 2000. Some biological characteristics of the brown trout Salmo trutta fario (Linnaeus, 1758) in Tonekabon River. Iranian Journal of Fisheries Sciences, 9(3):21-34. In Farsi.

Afraei, M. A. and Hassannia, M. R. 1999. Some biological aspects of the smallest goby fish (Knipowitschia caucasica Berg, 1916) in Gorgan Bay, Caspian Sea. Iranian Journal of Fisheries Sciences, 8(2):59-68, 5. In Farsi.

Afraei, M. and Lalooie, F. 2000. Fishes distribution assay in Tonekabon River. Iranian Journal of Fisheries Sciences, 9(1):1-14, 1. In Farsi.

Afraei Bandpyi, M. A., Salmani, A., Khoshbavar Rostami, H. R. and Paratkandeh Hagbighy, F. 2004. Biosystematic study of (Clupeidae) Alosa in Mazandaran and Golestan Province coastal waters. Iranian Journal of Fisheries Sciences, 13(3):13-26. In Farsi.

Afrasiabi, K. L. 2003. The environmental movement in Iran: perspectives from below and above. Middle East Journal, 57(3):432-448.

Afraz, A. 1996. Classification of Anzali Lagoon rivers. Iranian Fisheries Scientific Journal, 5(1):1-17, 1. In Farsi.

Agah, H., Leermakers, M., Elskens, M., Fatemi, S. M. R. and Baeyens, W. 2007. Total mercury and methyl mercury concentrations in fish from the Persian Gulf and the Caspian Sea. Water, Air, & Soil Pollution, 181(1-4):95-105.

Aghaei Moghadam, A. A. and Aslan Parviz, H. 2006. The study on the nutrition of the juvenile sturgeon (Acipenser persicus) in fish ponds of Shahid Rajaie's centre (1999). Pajouhesh va Sazandegi, 60:77-83. In Farsi.

Ahmad, M. F. and Niazi, M. S. 1988. Important Edible Fishes of Pakistan. Zoological Survey Department, Government of Pakistan, Karachi. ix + 31 pp.

Ahmad, T., Slaman (sic), N. A. and Hussain, N. A. 1983. Effects of some experimental conditions on the behaviour and survival of Liza abu (Heckel) from Basrah, Iraq. Journal of the Faculty of Marine Science, Jeddah, 3:111-117.

Ahmad, T. A. and Hussain, N. A. 1982. Observations on the food of young Liza abu (Heckel) from Salihyia river (Basrah, Iraq). Iraqi Journal of Marine Science, Basrah, 1(1):79-88.

Ahmad, T. A., Salman, N. A. and Hussain, N. A. 1985. Effect of some laboratory controlled conditions on the survival of Liza abu (Heckel). Kuwait Bulletin of Marine Science, 1985(6):125-131.

Ahmadi, M. R. 1988. The basic principles of breeding in fish culture. Journal of the Veterinary Faculty of the University of Tehran, 44(1):1-20 In Farsi.

Ahmadi, M. R. 1993. Aquaculture industry in Iran, p. 69-79. In: I Chiu Liao, Chung-Zen Shyu and Nai-Hsien Chao. (Eds.). Aquaculture in Asia. Proceedings of the 1990 Asian Productivity Organization Symposium on Aquaculture, Keelung, Taiwan, 5-13 September, 1990.

Ahmadi, M. R. and Alizadeh, M. 2004. Effects of dietary protein and energy levels on rainbow trout (Oncorhynchus mykiss) reared in brackish water. Iranian Journal of Fisheries Sciences, 4(1):77-88, 123.

Ahmadi, M. R. and Rezai, M. 1998. Comparison of feeding preference and growth of grass carp (Ctenopharyngodon idella) fed different plant with emphasis on chemical composition. Journal of the Faculty of Veterinary Medicine, University of Tehran, 53(1-2):?-5. In Farsi.

Ahmadi, M. R., Safi, Sh., Bjerkeng, B., Gerami, A., Ranaii Siadat, S. O. and Khodarahmi, R. 2005. Caspian Sea gammarus (Pontogammarus maeoticus) as a carotenoid source for muscle pigmentation of rainbow trout (Oncorhynchus mykiss). Iranian Journal of Fisheries Sciences, 5(1):1-12.

Ahmadi, M. R. und Wossughi, Gh. 1988. Ein Beitrag zur fischereilichen Bedeutung des Hamun-Sees, Iran. Österreichs Fischerei, 41(10):209-216.

Ahmed, A. H. and Taher, M. M. 1988. Observations on the extensive and intensive cultured carp, Cyprinus carpio L. in Basrah, Iraq. Basrah Journal of Agricultural Science, 1:35-47.

Ahmed, H. A. 1982. Crowth (sic) of the cyprinid fish, Barbus luteus (Heckel) in Tharthar Reservoir, Iraq. Bulletin of the Basrah Natural History Museum, 5:3-15.

Ahmed, H. A., Al-Mukhtar, M. A. and Al-Adhub, A. H. Y. 1984. The reproductive biology of Carasobarbus luteus (Pisces, Cyprinidae) in Al-Hammar Marsh, Iraq. Cybium, 8(4):69-80.

Ahmed, S. S., Linden, A. L. and Cech, J. J. 1988. A rating system and annotated bibliography for the selection of appropriate, indigenous fish species for mosquito and weed control. Bulletin of the Society for Vector Ecology, 13(1):1-59.

Ahnelt, H. 2003. The postcranial skeleton of the benthophiline gobiids Anatirostrum and Benthophilus (Teleostei: Gobiidae). Folia Zoologica, Prague, 52(2):213-221.

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Churchill (Consul). 1877. Persian fisheries. Forest and Stream, London, 9:249-250.

Ciepielewski, W., Martyniak, A. and Szczerbowski, J. A. 2001. Ichthyofauna in the Dokan and Derbendikhan reservoirs. Archives of Polish Fisheries, 9(supplement 1):157-170.

Čihar, J. 1975. Geographical and ecologic variability of perch [Perca fluviatilis (Linnaeus)] and history of its distribution from Eurasia to North America. Sborník Národního Muzea v Praze, 31B(1-2):57-89.

CIRSPE (Italian Research and Study Centre for the Fishery). 2006a. Aquaculture Development in Sistan-Baluchestan 2005-2008. Technical Report, The Hamun Lake, Preliminary Analysis. CIRSPE (Italian Research and Study Centre for the Fishery), Rome. 70 pp. (www.iranitaly.net).

CIRSPE (Italian Research and Study Centre for the Fishery). 2006b. Aquaculture Development in Sistan-Baluchestan 2005-2008. Technical Report, Artificial reproduction of Schizothorax zarudny (sic). Methodological approach, Zabol/Zahedan March-June 2006. CIRSPE (Italian Research and Study Centre for the Fishery), Rome. 48 pp., figs. (www.iranitaly.net).

CITES. 2001. CITES Identification Guide - Sturgeons and Paddlefishes. Guide to the Identification of Sturgeon and Paddlefish Species Controlled under the Convention on International Trade in Endangered Species of Wild Fauna and Flora. Environment Canada, Ottawa. Pagination various.

Civil, M. 1961. The home of the fish. A new Sumerian literary composition. Iraq, 23:154-175.

Clapp, D. F., Schneeberger, P. J., Jude, D. J., Madison, G. and Pistis, C. 2001. Monitoring round goby (Neogobius melanostomus) population expansion in eastern and northern Lake Michigan. Journal of Great Lakes Research, 27(3):335-341.

Clark, R. B. 1986. Marine Pollution. Clarendon Press, Oxford. xiii + 215 pp.

Clay, C. H. 1977. Report on a mission to Iran to investigate the potential for fish production from man-made lakes. Food and Agriculture Organization, Rome. MS. (not seen).

Clayton, D. A. and Vaughan, T. C. 1988. Ethogram of Boleophthalmus boddarti (Pallas)(Teleostei, Gobiidae), a mudskipper found on the mudflats of Kuwait. Journal of the University of Kuwait (Sciences), 15:115-138.

Clugston, J. P. and Shireman, J. V. 1987. Triploid grass carp for aquatic plant control. Fish and Wildlife Service, Washington, Fish and Wildlife Leaflet, 8:3 pp.

Coad, B. W. 1979a. Range extension for the snakehead Ophiocephalus gachua Hamilton-Buchanan (Osteichthyes: Channidae) in Iran. Journal of the Bombay Natural History Society, 75(2):(1978):500-501.

Coad, B. W. 1979b. Poisonous and venomous freshwater fishes of Iran. Pahlavi Medical Journal, 9(4):388-407.

Coad, B. W. 1980a. A provisional, annotated check-list of the freshwater fishes of Iran. Journal of the Bombay Natural History Society, 76(1)(1979):86-105.

Coad, B. W. 1980b. A re-description of Aphanius ginaonis (Holly, 1929) from southern Iran (Osteichthyes: Cyprinodontiformes). Journal of Natural History, 14(1):33-40.

Coad, B. W. 1980c. Environmental change and its impact on the freshwater fishes of Iran. Biological Conservation, 19(1):51-80.

Coad, B. W. 1981a. Glyptothorax silviae, a new species of sisorid catfish from southwestern Iran. Japanese Journal of Ichthyology, 27(4):291-295.

Coad, B. W. 1981b. First record of the milkfish, Chanos chanos (Forskal, 1775) from Iran and the Persian Gulf. Journal of the Bombay Natural History Society, 77(3)(1980):522-524.

Coad, B. W. 1981c. A new genus and species of cichlid endemic to southern Iran. Program of the American Society of Ichthyologists and Herpetologists, 61st Annual Meeting, 21-26 June 1981, Corvallis, Oregon. 1 p. (Abstract).

Coad, B. W. 1981d. Fishes of Afghanistan, an annotated checklist. Publications in Zoology, National Museums of Canada, 14:v + 26 pp.

Coad, B. W. 1981e. Pseudophoxinus persidis, a new cyprinid fish from Fars, southern Iran. Canadian Journal of Zoology, 59(11):2058-2063.

Coad, B. W. 1981f. A bibliography of the sticklebacks (Gasterosteidae: Osteichthyes). Syllogeus, Ottawa, 35:1-142.

Coad, B. W. 1982a. A new genus and species of cichlid endemic to southern Iran. Copeia, 1982(1):28-37.

Coad, B. W. 1982b. Review of "Euphrates and Tigris, Mesopotamian Ecology and Destiny" by Julian Rozska (sic) with contributions by J. F. Talling, F. R. S. and Dr. K. E. Bainster (sic). Monographiae Biologicae, Volume 38, Dr. W. Junk by (sic) Publishers, The Hague, 1980. x + 122 pp., 36 figs., map.-U.S. $33.31. Matsya, Bulletin of the Indian Society of Ichthyologists, 7(1981):102-104.

Coad, B. W. 1982c. Garra persica Berg, 1913, a valid species of cyprinid fish from southern Iran. Cybium, 6(2):97-100.

Coad, B. W. 1982d. The identity of Alburnus maculatus Keyserling, a cyprinid fish from Esfahan Province, Iran. Japanese Journal of Ichthyology, 29(2):227-228.

Coad, B. W. 1982e. Studies on the systematics and zoogeography of the freshwater fishes of Iran. Programme of the Fourth Congress of European Ichthyologists, 20.-24. 9. 1982, Hamburg, West Germany. 1 p. (Abstract).

Coad, B. W. 1982f. A re-description and generic re-assignment of Kosswigobarbus kosswigi (Ladiges, 1960), a cyprinid fish from Turkey and Iran. Mitteilungen aus dem hamburgischen Zoologischen Museum und Institut, 79:263-265.

Coad, B. W. 1983. Review of "The Freshwater Fishes of India, Pakistan, Bangladesh, Burma and Sri Lanka - A Handbook". By K. C. Jayaram. 1981. Zoological Survey of India, Calcutta. xxii + 475 pp., 208 figs., 13 plates (2 color). Rs 100.00 or $22.50. Copeia, 1983(1):280-282.

Coad, B. W. 1984. Acanthobrama centisquama Heckel and the validity of the genus Mirogrex Goren, Fishelson and Trewavas (Osteichthyes: Cyprinidae). Hydrobiologia, 109(3):275-278.

Coad, B. W. 1985. Alburnus doriae De Filippi, 1864 a synonym of Leuciscus lepidus (Heckel, 1843)(Osteichthyes: Cyprinidae). Matsya, 9-10(1983-1984):173-175.

Coad, B. W. 1987a. The tooth-carps of Iran. Program and Abstracts, American Society of Ichthyologists and Herpetologists, 67th Annual Meeting, 21-26 June 1987, Albany, New York. p. 39 (Abstract).

Coad, B. W. 1987b. Zoogeography of the Freshwater Fishes of Iran, pp. 213-228, 1 figure, 2 tables. In: Krupp, F., Schneider, W. and Kinzelbach, R. (Eds.). Proceedings of the Symposium on the Fauna and Zoogeography of the Middle East, Mainz, 1985. Beihefte zum Tübinger Atlas des Vorderen Orients, Reihe A (Naturwissenschaften), 28, Dr. Ludwig Reichert Verlag, Wiesbaden, 338 pp.

Coad, B. W. 1988. Aphanius vladykovi, a new species of tooth-carp from the Zagros Mountains of Iran (Osteichthyes: Cyprinodontidae). Environmental Biology of Fishes, 23(1-2):115-125. (Reprinted with same pagination in McAllister, D. E. and Kott, E. (Eds.). On lampreys and fishes: a memorial anthology in honor of Vadim D. Vladykov. Developments in Environmental Biology of Fishes, 8:1-162, Kluwer Academic Publishers, Dordrecht/Boston/London).

Coad, B. 1989. Tooth-carps and Tangs or Orogeny and Speciation in the Zagros Mountains of Iran. Research Results Conference, National Museum of Natural Sciences, Ottawa, 2-3 October 1989. 2 pp. (Abstract).

Coad, B. W. 1991a. The qanat ichthyofauna of Iran. Research Results CONFERENCE des résultats de recherche 1991, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, March 21, 1991 / le 21 mars 1991. p. 27 (Abstract).

Coad, B. W. 1991b. Fishes of the Tigris-Euphrates Basin: A Critical Checklist. Syllogeus, Ottawa, 68:1-49.

Coad, B. W. 1992a. Marine Fishes of the Persian Gulf and Sea of Oman. Checklist and Bibliography. Syllogeus, Ottawa, MS. 214 pp.

Coad, B. W. 1992b. Out of Africa? The large-scale barbs of Southwest Asia. Collections and Research CONFERENCE des collections et recherche 1992, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, February 18, 1992 / le 18 février 1992. p. 28. (Abstract).

Coad, B. W. 1992c. Zoogeography of the fishes of Mesopotamia. Collections and Research CONFERENCE des collections et recherche 1992, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, February 18, 1992 / le 18 février 1992. p. 30. (Abstract).

Coad, B. W. 1992d. Environmental change and its impact on the freshwater fishes of Iran, pp. 103-104. In: Trussell, D. (Ed.). The Social and Environmental Effects of Large Dams. Volume III. A Review of Literature. Wadebridge Ecological Centre, Camelford, U.K. xiv + 243 pp. (abstract of Coad (1980c)).

Coad, B. W. 1993. Expedition Field Techniques. Fishes. Expedition Advisory Centre, Royal Geographical Society, London. iv + 82 pp., 25 illustrations.

Coad, B. W. 1994a. Exotic and transplanted fishes in Southwest Asia. VIII Congress Societas Europaea Ichthyologorum, Oviedo, Spain, September 26 to October 2, 1994, p. 20 (abstract).

Coad, B. W. 1994b. Fishes from the qanats of Iran. VIII Congress Societas Europaea Ichthyologorum, Oviedo, Spain, September 26 to October 2, 1994, p. 75 (abstract).

Coad, B. W. 1994c. Criteria for assessing the conservation status of taxa. Science Forum Scientifique, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, November 21 Novembre 1994, p. 35 (abstract).

Coad, B. W. 1994d. Exotic and transplanted fishes in Southwest Asia. Science Forum Scientifique, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, November 21 Novembre 1994, p. 36 (abstract).

Coad, B. W. 1995a. Freshwater Fishes of Iran. Acta Scientiarum Naturalium Academiae Scientiarum Bohemicae, Brno, 29(1):1-64.

Coad, B. W. 1995b. Expedition Field Techniques: Fishes. Expedition Advisory Centre, Royal Geographical Society, London. 2nd Edition. v + 97 pp., 27 illustrations.

Coad, B. W. 1996a. Systematics of the tooth-carp genus Aphanius Nardo, 1827 (Actinopterygii: Cyprinodontidae) in Fars Province, southern Iran. Biologia, Bratislava, 51(2):163-172.

Coad, B. W. 1996b. Systematics of the shah mahi, Chalcalburnus chalcoides (Güldenstädt, 1772), in the southern Caspian Sea basin (Actinopterygii: Cyprinidae). Zoology in the Middle East, 12:65-70.

Coad, B. W. 1996c. Threatened fishes of the world: Iranocypris typhlops Bruun and Kaiser, 1944 (Cyprinidae). Environmental Biology of Fishes, 46(4):374.

Coad, B. W. 1996d. Freshwater fishes of Iranian and Pakistani Baluchistan. Second Symposium on Fish and Fisheries of Pakistan, November, 25, 26, 1996, Department of Zoology, Government College, Lahore, p. 25-26 (abstract).

Coad, B. W. 1996e. Zoogeography of the fishes of the Tigris-Euphrates Basin. Zoology in the Middle East, 13:51-70.

Coad, B. W. 1996f. Exotic fish species in the Tigris-Euphrates basin. Zoology in the Middle East, 13:71-83.

Coad, B. W. 1996g. Fishes from the qanats of Iran. Publicaciones Especiales Instituto Español de Oceanografía, 21:63-79.

Coad, B. W. 1996h. Exotic and transplanted fishes in Southwest Asia. Publicaciones Especiales Instituto Español de Oceanografía, 21:81-106.

Coad, B. W. 1997a. Status of shad in Iranian waters? Shad Journal, 2(2):3.

Coad, B. W. 1997b. Systematic biodiversity in the freshwater fishes of Iran. Ninth International Congress of European Ichthyologists (CEI9) "Fish Biodiversity". Italy 1997 (Napoli-Trieste). Book of Abstracts, p. 25.

Coad, B. W. 1997c. Freshwater fishes of Iranian and Pakistani Baluchistan. Biologia, Lahore, 42(1 & 2)(1996):1-18.

Coad, B. W. 1997d. Systematic biodiversity in the freshwater fishes of Iran. Science FORUM scientifique, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, November 20 novembre, 1997, p. 27 (abstract).

Coad, B. 1997e. Shad in Iranian waters. The Shad Journal, Seattle, 2(4):4-8.

Coad, B. W. 1998a. Threatened fishes of the world: Lebias ginaonis (Holly, 1929)(Cyprinodontidae). Environmental Biology of Fishes, 51(3):284.

Coad, B. W. 1998b. Lebias ginaonis (Holly, 1929). Holly's pupfish. Killi Kontakt, Wommelgem, 4 pp., 1 figure.

Coad, B. W. 1998c. Description of Barbus sublimus Coad and Najafpour, 1997, 2 pp., 2 figures. In: Berrebi, P. (Ed.). Atlas Barbus on the Internet, Université Montpellier, France (http://162.38.181.13/webbarbus/).

Coad, B. W. 1998d. Description of Barbus grypus Heckel, 1843, 2 pp., 2 figures. In: Berrebi, P. (Ed.). Atlas Barbus on the Internet, Université Montpellier, France (http://162.38.181.13/webbarbus/).

Coad, B. W. 1998e. Systematic biodiversity in the freshwater fishes of Iran. Italian Journal of Zoology, 65 (Supplement):101-108. (Proceedings of the Ninth Congress of European Ichthyologists (CEI-9) "Fish Biodiversity" organised in Naples at the University Federico II and held in Trieste - Italy, 24-30 August 1997).

Coad, B. W. 1998f. Expedition Field Techniques: Fishes. Expedition Advisory Centre, Royal Geographical Society, London. 2nd Revised Edition. v + 97 pp., 27 illustrations. ISBN 0-907649-71-8.

Coad, B. W. 1999a. Researchs and scientists. Brian W. Coad. Iran Nature and Wildlife Magazine, Farvadin 1378(3):2 pp., 2 figures. In Farsi and English.

Coad, B. W. 1999b. (Fish). i. Freshwater fishes, p. 655-669. In: Yarshater, E. (Ed.). Encyclopædia Iranica. Bibliotheca Persica Press, New York. Volume IX, Fascicle 6. Festivals VIII-Fish.

Coad, B. W. 1999c. Verspreiding van Aphanius-soorten in Iran. Killi-Kontakt, Wommelgem, 27(4):102-110. In Flemish.

Coad, B. W. 2000a. Criteria for assessing the conservation status of taxa (as applied to Iranian freshwater fishes). Biologia, Bratislava, 55(5):539-557.

Coad, B. W. 2000b. Distribution of Aphanius species in Iran. Journal of the American Killifish Association, 33(6):183-191.

Coad, B. W. 2000c. Aphanius ginaonis (Holly, 1929). Holly's pupfish. Journal of the American Killifish Association, 33(6):192-194.

Coad, B. W. 2003. Freshwater fishes, 15 pp., 20 figures, 2 tables. In: Yarshater E. (Ed.). Encyclopædia Iranica (Daneshnameh-ye Iranika). Volume IX, Fascicule 6. Festivals VIII - Fish. Bibliotheca Persica Press, New York. (on-line version of Coad (1999b) at http://www.iranica.com/articles/v9f6/v9f641a.html, downloaded 30 July 2003).

Coad, B. W. 2004. Review of "Caviar: The Strange History and Uncertain Future of the World’s Most Coveted Delicacy” by Inga Saffron. 2002. Broadway Books, New York. xv + 270 pp. U.S. $35.95. Canadian Field-Naturalist, 117(2)(2003):324-325.

Coad, B. W. 2006. Endemicity in the freshwater fishes of Iran. Iranian Journal of Animal Biosystematics, 1(1)(2005):1-13.

Coad, B. W. 2007. Review of “The Complete Fauna of Iran” by Eskandar Firouz. 2005. I. B. Tauris. London, New York. xiv + 322 pp. US$90.00. Canadian Field-Naturalist, 120(1)(2006):114-116.

Coad, B. W. 2008. Fishes of Tehran Province and adjacent areas. Shabpareh Publications, Tehran. 244 pp., 38 figures. ISBN 978-600-5038-02-6. (40,000 rials).

Coad, B. W. (Continuing). Bibliography on the Freshwater Fishes of Iran. http://gause.biology.ualberta.ca/Keivany/irbiblcoad.html. First posted 10 September 1996. Page originally maintained by Yazdan Keivany at the University of Alberta, Edmonton and now replaced by this website at www.briancoad.com, maintained by Brian W. Coad and Nicholas P. Coad, Ottawa, Ontario, Canada.

Coad, B. W. (Continuing). Freshwater Fishes of Iran. First posted 26 February 2002 at www.briancoad.com, maintained by Brian W. Coad and Nicholas P. Coad, Ottawa, Ontario, Canada.

Coad, B. W. (Continuing). Freshwater Fishes of Iraq. First posted 28 September 2005 at www.briancoad.com, maintained by Brian W. Coad and Nicholas P. Coad, Ottawa, Ontario, Canada.

Coad, B. W. and Abdoli, A. 1993a. Exotic fish species in the fresh waters of Iran. Science Forum scientifique, Canadian Museum of Nature / Musée canadien de la nature, Ottawa, November 17 novembre 1993. p. 40. (Abstract).

Coad, B. W. and Abdoli, A. 1993b. Exotic fish species in the fresh waters of Iran. Zoology in the Middle East, 9:65-80.

Coad, B. W. and Abdoli, A. 1996. Biodiversity of Iranian freshwater fishes. Abzeeyan, Tehran, 7(1):4-10, IV. In Farsi, English abstract.

Coad, B. W. and Abdoli, A. 2000a. Rhinogobius cf. similis Gill, 1859, a goby new to the fish fauna of Iran and the problem of alien invasions. Zoology in the Middle East, 20:55-59.

Coad, B. W. and Abdoli, A. 2000b. Systematics of an isolated population of tooth-carp from northern Iran (Actinopterygii: Cyprinodontidae). Zoology in the Middle East, 21:87-102.

Coad, B. W. and Al-Hassan, L. A. J. 1989a. A Bibliography of the Fishes of the Tigris-Euphrates Basin / Bibliographie der Fische des Euphrat-Tigris-Beckens. Max Kasparek Verlag, Heidelberg. 56 pp. ISBN 3-925064-05-2. DM 19.80. (dated 1988).

Coad, B. W. and Al-Hassan, L. A. J. 1989b. Freshwater shark attacks at Basrah, Iraq. Zoology in the Middle East, 3:49-53.

Coad, B. W., Alkahem, H. F. and Behnke, R. J. 1983. Acanthobrama hadiyahensis, a new species of cyprinid fish from Saudi Arabia. Publications in Natural Sciences, National Museums of Canada, 2:v + 6 pp.

Coad, B. W., Atz, J. W. and Keivany, Y. 2000. Fish imagery in art 82: Jonah and the fish. Environmental Biology of Fishes, 57(1):9.

Coad, B. W. and Delmastro, G. B. 1985. Notes on a sisorid catfish from the Black Sea drainage of Turkey. Cybium, 9(3):221-224.

Coad, B. W. and Holčík, J. 1999. Systematics of the cyprinid fish Chalcalburnus atropatenae (Berg, 1925) from the Lake Orumiyeh basin in northwest Iran. Biologia, Bratislava, 54(2):179-186.

Coad, B. W. and Holčík, J. 2000. On Silurus species from Iran (Actinopterygii: Siluridae). Folia Zoologica, Prague, 49(2):139-148.

Coad, B. W. and Hussain, N. A. 2007. First record of the exotic species Hemiculter leucisculus (Actinopterygii: Cyprinidae) in Iraq. Zoology in the Middle East, 40:107-109.

Coad, B. W., Hussain, N. A., Ali, T. S. and Limburg, K. E. 2003. Middle Eastern shads. In: Limburg, K. E. and Waldman, J. R. (Eds.). Biodiversity, Status, and Conservation of the World’s Shads. American Fisheries Society Symposium, 35:59-67 (ISBN 1-888569-51-4, ISSN 0892-2284).

Coad, B. W. and Keivany, Y. 1998b. Lebias vladykovi (Coad, 1988). Killi-Kontakt, Wommelgem, 26(3):43-48. In Flemish.

Coad, B. W. and Keivany, Y. 2000. Aphanius vladykovi Coad, 1988. Zagros pupfish, mahi-e gour-e khari. Journal of the American Killifish Association, 33(6):195-198.

Coad, B. W. and Keivany, Y. 2002a. Aphanius vladykovi Coad, 1988 "Zagros pubfish" (sic) mahi-e gour-e khari. Skandinaviska Killi Sällskapet Killibladet, 2002(2):3-6, 1 figure. In Swedish (translation of Coad and Keivany, 2000).

Coad, B. W. and Keivany, Y. 2002b. Reviews of "Atlas of Iranian Fishes: Gilan Inland Waters. K. Abbasi, A. Valipour, D. Talebi Haghighi, A. Sarpanah, and Sh. Nezami. 1999. Gilan Fisheries Research Centre, Rasht. vi + 113 p. 35,000 Rls ($4.38)(paperback). The Inland Water Fishes of Iran. A. Abdoli. 2000. Iranian Museum of Nature and Wildlife, Tehran. ISBN: 964-6902-01-4. 378 p. 75,000 Rls ($9.38)(hardbound). A Guide to the Fauna of Iran. E. Firouz. 2000. Iran University Press (University Publication Centre), Tehran. ISBN: 964-01-0956-8. vi + 491 p. 45,000 Rls ($5.63)(hardbound). Freshwater Fishes of Iran. H. Mohammadian. 1999. Sepehr Publications Centre, Tehran. ISBN: 946-6123-18.X. vii + 178 p. 12,000 Rls ($1.50)(paperback)". Copeia, 2002(4):1164-1166.

Coad, B. W. and Keyzer-de Ville, N. 2004. On the systematics and distribution of the snow trout, Schizothorax pelzami Kessler, 1870, in Iran (Actinopterygii: Cyprinidae). Zoology in the Middle East, 32:57-62.

Coad, B. W. and Keyzer-de Ville, N. 2005. On the validity of the species in the snow-trout genus Schizocypris Regan, 1914 (Cyprinidae: Actinopterygii). Zoology in the Middle East, 35:35-42.

Coad, B. W. and Krupp, F. 1983. Redescription of Barilius mesopotamicus Berg, 1932 a poorly known cyprinid fish from the Tigris-Euphrates basin. Cybium, 7(1):47-56.

Coad, B. W. and Krupp, F. 1994. Capoeta aculeata (Valenciennes in Cuv. & Val., 1844), a valid species of cyprinid fish from Iran (Teleostei: Cyprinidae). Zoology in the Middle East, 10:63-72.

Coad, B. W. und Kuru, M. 1986. Bibliographie der Fische der Türkei/A Bibliography of the Fishes of Turkey, pp. 15-77. In: Kasparek, M. (Ed.). Zoologische Bibliographie der Türkei. Zoological Bibliography of Turkey. Pisces, Amphibia, Reptilia. Max Kasparek Verlag, Heidelberg. 118 pp. ISBN 3-925064-01-X. DM 24.50.

Coad, B. W. and McAllister, D. E. (Continuing). Dictionary of Ichthyology. First posted 4 September 2002 at www.briancoad.com, maintained by Brian W. Coad and Nicholas P. Coad, Pure Throttle Technologies Inc., Ottawa, Ontario.

Coad, B. W. and Najafpour, N. 1997. Barbus sublimus, a new species of cyprinid fish from Khuzestan Province, Iran. Ichthyological Exploration of Freshwaters, 7(3):273-278.

Coad, B. W. and Nalbant, T. T. 2005. A new genus and a new species of a remarkable nemacheilid fish from Iran (Pisces: Ostariophysi: Nemacheilidae). Travaux du Muséum National d’Histoire Naturelle "Grigore Antipa", Bucharest, 48:303-308.

Coad B. W. and Nalbant T. T. 2006. On the systematic position of the recently described genus and species Ilamnemacheilus longipinnis from Iran (Pisces: Ostariophysi: Nemacheilidae). 3rd International Conference Loaches of the Genus Cobitis and Related Genera. Biology, Systematics, Genetics, Distribution, Ecology, Conservation. Sibenik, Croatia, 24-29 September 2006 (title).

Coad, B. W. and Papahn, F. 1988. Shark attacks in the rivers of southern Iran. Environmental Biology of Fishes, 23(1-2):131-134. (Reprinted with same pagination in McAllister, D. E. and Kott, E. (Eds.). On lampreys and fishes: a memorial anthology in honor of Vadim D. Vladykov. Developments in Environmental Biology of Fishes, 8:1-162, Kluwer Academic Publishers, Dordrecht/Boston/London).

Coad, B. W. and Sarieyyüpoğlu, M. 1988. Cobitis elazigensis, a new species of cobitidid fish from Anatolia, Turkey. Japanese Journal of Ichthyology, 34(4):426-430.

Coad, B. W. and Vilenkin, B. Ya. 2004. Co-occurrence and zoogeography of the freshwater fishes of Iran. Zoology in the Middle East, 31:53-61.

Coad, B. W. with Waszczuk, H. and Labignan, I. 1995. Encyclopedia of Canadian Fishes. Canadian Museum of Nature, Ottawa and Canadian Sportfishing Productions, Waterdown. viii + 928 pp.

COFREPECHE. 1997. Iranian Fisheries Sector Study. Parts 1-4. Report prepared for Shilat (Fisheries Department), Jihad-e Sazandagi, Islamic Republic of Iran.

Collares-Pereira, M. J. 1994. The karyology of barbins and the possible plesiomorphic condition of polyploïdy in Cyprinidae. Bulletin français de la pêche et de la pisciculture, 334:191-199.

Collette, B. B. 1982. Rediscovery of Hyporhamphus xanthopterus, a half beak endemic to Vembanad Lake, Kerala, southern India. Matsya, 7(1981):29-40.

Collette, B. B., Ali, M. A., Hokanson, K. E. F., Nagieć, M., Smirnov, S. A., Thorpe, J. E., Weatherly, A. H. and Willemsen, J. 1977. Biology of Percids. Journal of the Fisheries Research Board of Canada, 34:1890-1899.

Collette, B. B. and Bănărescu, P. 1977. Systematics and zoogeography of the fishes of the Family Percidae. Journal of the Fisheries Research Board of Canada, 34:1450-1463.

Compagno, L. J. V. 1984. FAO Species Catalogue. Volume 4. Sharks of the World. An Annotated and Illustrated Catalogue of Shark Species Known to Date. Part 1 - Hexanchiformes to Lamniformes, Part 2 - Carcharhiniformes. Food and Agriculture Organization, Rome, Fisheries Synopsis, 125(4), part 1:viii + 1-249; part 2:x + 251-655.

Compagno, L. J. V. 1988. Sharks of the Order Carcharhiniformes. Princeton University Press, Princeton, New Jersey. xxii + 486 pp., 21 figs., 35 plates.

Conrad, G. et Conrad, J. 1970. L'évolution quaternaire de la dépression du Lout (Iran oriental). Comptes Rendus de l'Académie des Sciences, Paris, 270, série D, 9:1672-1674.

Cook, J. M. 1985. The rise of the Achaemenids and establishment of their empire, pp. 200-291. In: Gershevitch, I. (Ed.). The Cambridge History of Iran. Volume 2. The Median and Achaemenian Periods. Cambridge University Press, Cambridge. 946 pp.

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